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''Pterophylla racemosa'', commonly known as the kāmahi, is an evergreen tree
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to New Zealand. It is a member of the
Cunoniaceae Cunoniaceae is a family of 27 genera and about 335 species of woody plants in the order Oxalidales, mostly found in the tropical and wet temperate regions of the Southern Hemisphere. The greatest diversity of genera are in Australia and Tasmania ...
family of plants, and it is often referred to by its former botanical name ''Weinmannia racemosa.'' Most members of the Cunoniaceae are native to tropical wet parts of the Southern Hemisphere, with the majority of ''Pterophylla'' species being native to either
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
and
Papuasia Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia. It comprises th ...
or to
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
; however, two ''Pterophylla'' species are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to New Zealand: ''Pterophylla racemosa'', and '' Pterophylla sylvicola''.'''' Whilst ''P. sylvicola'' is restricted to subtropical forests in the far north of
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
, ''P. racemosa'' is found growing across vast areas of the country as far south as
Stewart Island Stewart Island (, ' glowing skies', officially Stewart Island / Rakiura, formerly New Leinster) is New Zealand's third-largest island, located south of the South Island, across Foveaux Strait. It is a roughly triangular island with a la ...
and is found in a variety of climatic conditions from mild coastal areas to mountainous (sub)-alpine areas inland. The abundance of the kāmahi (''P. racemosa'') across New Zealand has led to it long being described as the common tree in the country.


Description

A very common tree throughout New Zealand, kāmahi is evergreen that grows up to 25 m in height (rarely higher) with a trunk up to 24 cm (Wardle & MacRae, 1966). The tree has dark green leathery leaves approximately 7.5 cm long and 4 cm wide. It produces masses of creamy flowers between October and March which have a sweet, scented smell. The flowers form in erect spikes like clusters and are highly attractive to a range of insects and birds.


Distribution and habitat

Kāmahi is native to New Zealand although the genus ''Pterophylla'' has a more widespread distribution in Madagascar,
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
, and the southwest Pacific. Most species of this genus are tropical or sub-tropical. In New Zealand, however, kāmahi is found south in colder climate such as alpine terrain. Kāmahi appears in lowland, montane, and subalpine forests and shrubland from central North Island south to
Stewart Island Stewart Island (, ' glowing skies', officially Stewart Island / Rakiura, formerly New Leinster) is New Zealand's third-largest island, located south of the South Island, across Foveaux Strait. It is a roughly triangular island with a la ...
. Kāmahi’s habitat is coastal to subalpine. It is very commonly widespread in coastal, lowland and montane forest, often becoming dominant in higher elevation montane forest in the higher areas of the North Island and western South Island.


Life cycle

Kāmahi is durable, hardy, and persistent meaning it will grow in most situations. Kāmahi often starts life as a dense shrub; however, given the space, it can develop into a hulking tree that grows erratically. The white candle-like flowers that bloom in spring open to release fine seeds to be blown in the wind. Spreading their seed easily in spring. It is a slow growing plant that takes time to establish roots, however, once established, they will grow to 25 m at their best.


Ecology

Kāmahi prefers well drained sites and tolerates infertile soil reasonably well. It is found on hillslopes, roadside cuttings and can even start out life perched on the trunks of tree ferns. It has medium water requirements, preferring the soil to become dry between waterings. Kāmahi enjoys free draining soil which is suited to silt and sand-based soil, although it is accepted that this tree does not necessarily need particular climates and it is an adaptable plant that grows in diverse areas. Kāmahi flowers in spring and bees use its nectar for honey. Plenty of insects and other invertebrates use the trees to feed off and move around. A study in the Ōrongorongo Valley forest near Wellington called “Invertebrate fauna of four tree species in Ōrongorongo valley, New Zealand, as revealed by truck traps” discovered that tree trunks provide an important pathway (especially at night) for many ground-living invertebrates that move from the forest floor to feed or breed on tree stems and in the canopy. The Department of Conservation are monitoring a kāmahi forest on the Kaitake Range. This investigation brings up the problem of possums and the damage they do to kāmahi trees. Their results showed how possums were damaging the trees from the canopy to the ground floor. This shows that one of the threats to kāmahi are introduced species. Kāmahi have not adapted and have very little protection against invaders. However with kāmahi being a very common tree throughout New Zealand, possums should not pose much of a threat to the species.


Fruit

The fruit is about 5mm in diameter. The berries are not edible according to Crowe, the main authority on New Zealand edibles. He said the first error probably started with the confusion with tawa, which has a similar Māori name. He does not, however, say it is actually poisonous and it has not found out if it is anywhere (Nicholls, 2000). It was, however, very valuable to Māori for other uses (see below).


In human culture

Before the arrival of Europeans, kāmahi was valuable to Māori and because of this, it was protected by tapu (spiritual restriction, means that kāmahi was almost sacred to Māori). Back when the land was being transformed by Māori and being set up for cultivation, there are legends that say Māori were careful not to cut down all the tree’s limbs, if they did they or their spouse may suffer unfortunate consequences. Kāmahi bark is a rich source of tannins and a reddish, permanent dye can be obtained by boiling and soaking the bark. It was historically used by some Māori to dye cloaks and mats and was considered to have a preservative effect on fishing lines. Following the arrival of Europeans to New Zealand, Kāmahi-based dyes were exported during the 1800s.Buchanan J. (1868). Sketch of the botany of Otago. ''Transactions of the New Zealand Institute 1: 22-53''


Medical uses

Inner bark is a purgative decoction. "Bark from west side of the tree, from which the outer rind has been scraped off, is steeped in hot water and the decoction taken internally as an aperient in cases of abdominal and thoracic pain". Bark infused in boiling water is a good tonic. Inner bark is also a laxative.


Gallery

Weinmannia racemosa L.f. (AM AK184482).jpg, Herbarium specimen WeinmanniaRacemosa.jpg, Botanical illustration Weinmannia racemosa in Paparoa National Park.jpg, Trunk Weinmannia racemosa flowers.jpg, Flowers Weinmannia racemosa kz5.jpg, ''Pterophylla racemosa'' tree in Haast


References

{{Subject bar, auto=y, d=y Trees of New Zealand racemosa Trees of mild maritime climate Endemic flora of New Zealand