Kyphosus Bigibbus
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''Kyphosus bigibbus'', the brown chub, grey drummer, darkfin drummer, insular rudderfish, grey chub, grey sea chub, southern drummer or topsail drummer is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a
sea chub The sea chubs, also known as rudderfish and pilot fish and in Hawaiian as ''enenue'' or ''nenue'', are a family, Kyphosidae, of fishes in the order Perciformes native to the Atlantic, Indian and Pacific Oceans usually close to shore in marine w ...
from the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Kyphosidae The sea chubs, also known as rudderfish and pilot fish and in Hawaiian as ''enenue'' or ''nenue'', are a family, Kyphosidae, of fishes in the order Perciformes native to the Atlantic, Indian and Pacific Oceans usually close to shore in marine w ...
. It is a herbivorous species which is found in subtropical and tropical seas worldwide.


Description

''Kyphosus bigibbus'' has an oval shaped body which is laterally compressed with a small head, a pointed snout and a slightly bulging forehead. The mouth is small and when closed the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
are hidden beneath the preorbital bones. The mouth is terminal and is almost oblique. The teeth are fixed and incisiform with their bases positioned horizontally in mouth, they have rounded crowns and have a curved, J shape. There are teeth are found on the centre of the roof of the mouth and on the tongue. The
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
is a similar height along its length The anterior part of the dorsal fin has 11 spines which fold down into a scaled furrow while the posterior part contains 10-12 rays. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has three spines and 10-12 moderately long soft rays and is reasonably long at its base. The symmetrical
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is forked but not extremely so. The
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
has 61-76 scales of which 51-60 are pored. The colour varies from uniform grey to brownish, silvery ventrally, bronze to greenish tint dorsally. There are very indistinct dark stripes on the body and there are usually whitish streaks on the cheek. The dorsal and anal fins are frequently dark with black margins on their soft-rayed parts. There is sometimes a dark patch visible on the lower posterior angle of the base of the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
. The caudal fin is usually dark. There are infrequent records of completely yellow individuals and very rate records of
albino Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...
s. The maximum
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of this species is and it has been weighed at .


Distribution

''Kyphosus bigibbus'' has a wide distribution in the Indian, Pacific and Atlantic Oceans. In the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
it has been recorded off
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
as far south as
Rottnest Island Rottnest Island ( nys, Wadjemup), often colloquially referred to as "Rotto", is a island off the coast of Western Australia, located west of Fremantle. A sandy, low-lying island formed on a base of aeolianite limestone, Rottnest is an A-class ...
, in eastern Australia it is found from southern
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
and
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
as far south as Montague Island, it occurs off northeastern New Zealand. It is also found in the
Coral Sea The Coral Sea () is a marginal sea of the South Pacific off the northeast coast of Australia, and classified as an interim Australian bioregion. The Coral Sea extends down the Australian northeast coast. Most of it is protected by the Fre ...
off
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
,
Lord Howe Island Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland P ...
,
Norfolk Island Norfolk Island (, ; Norfuk: ''Norf'k Ailen'') is an external territory of Australia located in the Pacific Ocean between New Zealand and New Caledonia, directly east of Australia's Evans Head and about from Lord Howe Island. Together with ...
and the
Kermadec Islands The Kermadec Islands ( mi, Rangitāhua) are a subtropical island arc in the South Pacific Ocean northeast of New Zealand's North Island, and a similar distance southwest of Tonga. The islands are part of New Zealand. They are in total are ...
. In the Indian Ocean it is thought to probably range widely along the northern coasts from Myanmar to Yemen. In the western Indian Ocean it occurs as far north is the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
and from the
Gulf of Aden The Gulf of Aden ( ar, خليج عدن, so, Gacanka Cadmeed 𐒅𐒖𐒐𐒕𐒌 𐒋𐒖𐒆𐒗𐒒) is a deepwater gulf of the Indian Ocean between Yemen to the north, the Arabian Sea to the east, Djibouti to the west, and the Guardafui Channe ...
off Somalia south to Madagascar,
Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...
. Its range then extends around the
Cape of Good Hope The Cape of Good Hope ( af, Kaap die Goeie Hoop ) ;''Kaap'' in isolation: pt, Cabo da Boa Esperança is a rocky headland on the Atlantic coast of the Cape Peninsula in South Africa. A common misconception is that the Cape of Good Hope is t ...
. It has not been confirmed to be present in the
Persian Gulf The Persian Gulf ( fa, خلیج فارس, translit=xalij-e fârs, lit=Gulf of Persis, Fars, ), sometimes called the ( ar, اَلْخَلِيْجُ ٱلْعَرَبِيُّ, Al-Khalīj al-ˁArabī), is a Mediterranean sea (oceanography), me ...
. In the western pacific it is found off southern Japan, Korea, and China as far south as Taiwan, but it is absent from equatorial areas. In the western Atlantic it has been observed off
Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...
,
Grand Cayman Island Grand Cayman is the largest of the three Cayman Islands and the location of the territory's capital, George Town. In relation to the other two Cayman Islands, it is approximately 75 miles (121 km) southwest of Little Cayman and 90 miles (1 ...
, Belize, and San Blas, Panama. In the eastern Atlantic it has been recorded from
Saint Helena Saint Helena () is a British overseas territory located in the South Atlantic Ocean. It is a remote volcanic tropical island west of the coast of south-western Africa, and east of Rio de Janeiro in South America. It is one of three constitu ...
,
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
,
Ascension Island Ascension Island is an isolated volcanic island, 7°56′ south of the Equator in the South Atlantic Ocean. It is about from the coast of Africa and from the coast of South America. It is governed as part of the British Overseas Territory o ...
, and probably occurs around the
Gulf of Guinea The Gulf of Guinea is the northeasternmost part of the tropical Atlantic Ocean from Cape Lopez in Gabon, north and west to Cape Palmas in Liberia. The intersection of the Equator and Prime Meridian (zero degrees latitude and longitude) is in the ...
Islands and along the west African coast to South Africa. It is also found along Atlantic coast of northwestern Africa as far north as Portugal, but it has not been recorded from the Mediterranean. It has been recorded from Brazil.


Habitat and biology

''Kyphosus bigibbus'' is found in shallow coral and rocky reefs, including exposed isolated, offshore reefs and around islands. They are frequently recorded in the surge zone, down to depths of . They are often recorded in mixed schools with other related species. The juveniles often hide among
flotsam In maritime law, flotsam'','' jetsam'','' lagan'','' and derelict are specific kinds of shipwreck. The words have specific nautical meanings, with legal consequences in the law of admiralty and marine salvage. A shipwreck is defined as the rema ...
and can disperse for long distances among this floating material. It is a herbivore which feeds mainly on algae of the genera ''
Sargassum ''Sargassum'' is a genus of brown (class Phaeophyceae) macroalgae (seaweed) in the order Fucales. Numerous species are distributed throughout the temperate and tropical oceans of the world, where they generally inhabit shallow water and coral re ...
'' and '' Turbinaria''.


Fisheries

''Kyphosus bigibbus'' is not targeted by fisheries over much of its distribution but where it is a fisheries quarry it is valued as a foodfish.


Species description

''Kyphosus bigibbus'' was first formally described by Bernard Germain de Lacépède in 1801 in volume 3 of ''Histoire naturelle des poissons'', Lacépède did not give a type locality but it is stated elsewhere as Fort Dauphin in Madagascar.


References


External links

* photo o
Kyphosus bigibbus
in iNaturalist * {{Taxonbar, from=Q2364096 bigibbus Fish described in 1801 Taxa named by Bernard Germain de Lacépède