The kathala croaker (''Kathala axillaris'') is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Sciaenidae Sciaenidae are a family of fish in the order Acanthuriformes. They are commonly called drums or croakers in reference to the repetitive throbbing or drumming sounds they make. The family consists of about 286 to 298 species in about 66 to 70 gen ...

, the drums and croakers. This species is found in the Indian Ocean off

South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...

. It is the only species in the

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

''Kathala''.

Taxonomy

The kathala croaker was first formally described as ''Corvina axillaris'' in 1830 by the French

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...

with its type locality given as

Malabar Malabar may refer to the following: People * Malabars, people originating from the Malabar region of India * Malbars or Malabars, people of Tamil origin in Réunion Places * Malabar Coast, or Malabar, a region of the southwestern shoreline o ...

in India. In 1969 R. S. Lal Mohan proposed the monospecific genus ''Kathala'' for this species. This taxon has been placed in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Cynoscioninae by some workers, but the 5th edition of ''

Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...

'' does not recognise subfamilies within the Sciaenidae which it places in the

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

Acanthuriformes Acanthuriformes is an order of ray-finned fishes, part of the Percomorpha clade. Some authorities place the fishes in the order within the Acanthuriformes in the suborders Acanthuroidea and Percoidea of the order Perciformes. Classification The ...

Etymology

The kathala croaker's genus name, was not explained by Lal Mohan but the name was recorded in 1801 by Patrick Russell in his ''Two Hundred Fishes: Collected at Vizagapatam on the Coast of Coromandel'' as a local name in India for this species and other members of the Sciaenidae. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''axillaris'' refers to a black mark over the base of the

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

Description

The kathala croaker has a body which has a depth which is almost equal to the length of its head. The

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

may be convex or truncate and has rounded corners. It has a carrot-shaped swimbladder with a pair of simple appendages at its head end. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

is divided by a deep incision, forward of the incision it is supported by 10 spines and to the rear of the incision there is a single spine and between 26 and 28 soft rays. The

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

is supported by 2 spines and 7 soft rays while the pectoral fins contain 17 or 18 rays. The

standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...

is 2.8 to 3.3 times the body's depth. There are small,

cycloid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...

s on the head and similarly sized cteniod scales on the body. The back is greyish-gree and the sides are slivery-yellow. The anterior part of the dorsal fin is black and tere is a black blotch above the base of the pectoral fin. This species has a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of , although is more typical.

Distribution and habitat

The kathala croakeris found in the northern Indian Ocean where it occurs along the coasts of Pakistan, Inida, Sri Lanka, Bangladesh and Myanmar. It is found at depths down to in shallow coastal and estuarine waters with substrates of mixed sand and mud.

References

{{Taxonbar, from1=Q3563738, from2=Q1986750 Fish described in 1830 Sciaenidae Taxa named by Georges Cuvier