Jackals are medium-sized

canids Canidae (; from Latin, ''canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within ...

native to

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

and

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed jackal (''Lupulella mesomelas'') and

side-striped jackal The side-striped jackal (''Lupulella adusta)'' is a canine native to central and southern Africa. Unlike the smaller and related black-backed jackal (''Lupulella mesomelas''), which dwells in open plains, the side-striped jackal primarily dwell ...

(''Lupulella adusta'') of sub-Saharan-Africa, and the

golden jackal The golden jackal (''Canis aureus''), also called common jackal, is a wolf-like canid that is native to Southeast Europe, Southwest Asia, South Asia, and regions of Southeast Asia. The golden jackal's coat varies in color from a pale creamy y ...

(''Canis aureus'') of south-central Europe and Asia. The African

golden wolf The African wolf (''Canis lupaster'') or golden wolf, formerly known as the African golden jackal, is a canine native to North Africa, West Africa, the Sahel, northern East Africa, and the Horn of Africa. It is listed as least concern on the ...

(''Canis lupaster'') was also formerly considered as a jackal. While they do not form a

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

, all jackals are opportunistic omnivores, predators of small to medium-sized animals and proficient

scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...

s. Their long legs and curved

canine teeth In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened howeve ...

are adapted for hunting small

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s, birds, and

reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...

s, and their large feet and fused leg bones give them a physique well-suited for long-distance running, capable of maintaining speeds of for extended periods of time. Jackals are

crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...

, most active at dawn and dusk. Their most common

social unit The term "level of analysis" is used in the social sciences to point to the location, size, or scale of a research target. "Level of analysis" is distinct from the term " unit of observation" in that the former refers to a more or less integrated ...

is a

monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...

pair, which defends its territory from other pairs by vigorously chasing intruding rivals and marking landmarks around the

territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...

with their

urine Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excretion, excreted from the body through the urethra. Cel ...

and

feces Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...

. The territory may be large enough to hold some young adults, which stay with their parents until they establish their own territories. Jackals may occasionally assemble in small packs, for example, to scavenge a carcass, but they normally hunt either alone or in pairs.

Etymology

The English word "jackal" dates back to 1600 and derives from the French ''chacal'', from Turkish çakal, derived from the

Persian Persian may refer to: * People and things from Iran, historically called ''Persia'' in the English language ** Persians, the majority ethnic group in Iran, not to be conflated with the Iranic peoples ** Persian language, an Iranian language of the ...

', which is in turn derived from the

Sanskrit Sanskrit (; attributively , ; nominally , , ) is a classical language belonging to the Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had diffused there from the northwest in the late ...

शृगाल ''śṛgāla'' meaning "the howler".

Taxonomy and relationships

Similarities between jackals and coyotes led

Lorenz Oken Lorenz Oken (1 August 1779 – 11 August 1851) was a German naturalist, botanist, biologist, and ornithologist. Oken was born Lorenz Okenfuss (german: Okenfuß) in Bohlsbach (now part of Offenburg), Ortenau, Baden, and studied natural history and ...

, in the third volume of his ''Lehrbuch der Naturgeschichte'' (1815), to place these species into a new separate genus, ''Thos'', named after the classical Greek word "jackal", but his theory had little immediate impact on taxonomy at the time. Angel Cabrera, in his 1932 monograph on the mammals of

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

, questioned whether or not the presence of a cingulum on the upper

molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

of the jackals and its corresponding absence in the rest of ''Canis'' could justify a subdivision of that genus. In practice, Cabrera chose the undivided-genus alternative and referred to the jackals as ''Canis'' instead of ''Thos''. Oken's ''Thos'' theory was revived in 1914 by

Edmund Heller Edmund Heller (May 21, 1875 – July 18, 1939) was an American zoologist. He was President of the Association of Zoos & Aquariums for two terms, from 1935-1936 and 1937-1938. Early life While at Stanford University, he collected specimens in th ...

, who embraced the separate genus theory. Heller's names and the designations he gave to various jackal species and subspecies live on in current taxonomy, although the genus has been changed from ''Thos'' to ''Canis''.Thos vs Canis
The wolf-like canids are a group of large carnivores that are genetically closely related because they all have 78

chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...

s. The group includes genus ''Canis'', ''Cuon'', and ''Lycaon''. The members are the

dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the wolf. Also called the domestic dog, it is derived from the extinct Pleistocene wolf, and the modern wolf is the dog's nearest living relative. Do ...

''(C. lupus familiaris)'',

gray wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly ...

(''C. lupus''),

coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...

(''C. latrans''),

(''C. aureus''),

Ethiopian wolf The Ethiopian wolf (''Canis simensis''), also called the Simien jackal and Simien fox, is a canine native to the Ethiopian Highlands. In southeastern Ethiopia it is also known as the horse jackal. It is similar to the coyote in size and build, ...

(''C. simensis''), black-backed jackal (''C. mesomelas''),

(''C. adustus''),

dhole The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It ...

(''Cuon alpinus''), and

African wild dog The African wild dog (''Lycaon pictus''), also called the painted dog or Cape hunting dog, is a wild canine which is a native species to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Ly ...

(''Lycaon pictus''). The latest recognized member is the African wolf (''C. lupaster''), which was once thought to be an African branch of the golden jackal. As they possess 78 chromosomes, all members of the genus ''

Canis ''Canis'' is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and den ...

'' are karyologically indistinguishable from each other, and from the dhole and the African hunting dog. The two African jackals are shown to be the most basal members of this clade, indicating the clade's origin from Africa. ''

Canis arnensis ''Canis arnensis'', the Arno River dog, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. The Arno River dog has been described as a small jackal-like dog. Its anatomy and morphology relate i ...

'' arrived in Mediterranean Europe 1.9 million years ago and is probably the ancestor of modern jackals. The paraphyletic nature of ''Canis'' with respect to '' Lycaon'' and ''

Cuon The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. ...

'' has led to suggestions that the two African jackals should be assigned to different genera, ''Schaeffia'' for the side-striped jackal and ''Lupulella'' for the black-backed jackal or ''Lupulella'' for both. The intermediate size and shape of the Ethiopian wolf has at times led it to be regarded as a jackal, thus it has also been called the "red jackal" or the "Simien jackal".

Species

Folklore and literature

fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...

es and coyotes, jackals are often depicted as clever sorcerers in the myths and legends of their regions. They are mentioned roughly 14 times in the

Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of texts of a ...

. It is frequently used as a literary device to illustrate desolation, loneliness, and abandonment, with reference to its habit of living in the ruins of former cities and other areas abandoned by humans. It is called "wild dog" in several translations of the Bible. In the

King James Bible The King James Version (KJV), also the King James Bible (KJB) and the Authorized Version, is an Bible translations into English, English translation of the Christian Bible for the Church of England, which was commissioned in 1604 and publis ...

, Isaiah 13:21 refers to 'doleful creatures', which some commentators suggest are either jackals or

hyenas Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the cla ...

. In the Indian '' Panchatantra'' stories, the jackal is mentioned as wily and wise. In Bengali tantrik tradition, they represent the goddess

Kali Kali (; sa, काली, ), also referred to as Mahakali, Bhadrakali, and Kalika ( sa, कालिका), is a Hinduism, Hindu goddess who is considered to be the goddess of ultimate power, time, destruction and change in Shaktism. In t ...

. It is said she appears as jackals when meat is offered to her. The

Serer religion The Serer religion, or ''a ƭat Roog'' ("the way of the Divine"), is the original religious beliefs, practices, and teachings of the Serer people of Senegal in West Africa. The Serer religion believes in a universal supreme deity called Roog ...

and

creation myth A creation myth (or cosmogonic myth) is a symbolic narrative of how the world began and how people first came to inhabit it., "Creation myths are symbolic stories describing how the universe and its inhabitants came to be. Creation myths develop ...

posits jackals were among the first animals created by

Roog Roog or Rog (Koox in the Cangin languages) is the Supreme God and creator of the Serer religion of the Senegambia region. Thiaw, Issa Laye, "La Religiosite de Seereer, Avant et pendant leur Islamisation". Ethiopiques no: 54, Revue semestrielle ...

, the supreme deity of the

Serer people The Serer people are a West African ethnoreligious group.

Etymology

Taxonomy and relationships

Species

Folklore and literature

References

Further reading

External links