''Yucca brevifolia'' is a plant species belonging to the genus ''
Yucca
''Yucca'' is a genus of perennial shrubs and trees in the family Asparagaceae, subfamily Agavoideae. Its 40–50 species are notable for their rosettes of evergreen, tough, sword-shaped leaves and large terminal panicles of white or whitish ...
''. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca.
This
monocotyledon
Monocotyledons (), commonly referred to as monocots, ( Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
ous
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
is native to the arid
Southwestern United States
The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, N ...
, specifically
California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
,
Arizona
Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a U.S. state, state in the Southwestern United States. It is the list of U.S. states and territories by area, 6th largest and the list of U.S. states and territories by population, 14 ...
,
Utah
Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to its ...
, and
Nevada
Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
, and to northwestern
Mexico
Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
It is confined mostly to the
Mojave Desert
The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily i ...
between elevation. It thrives in the open grasslands of Queen Valley and
Lost Horse Valley
Lost Horse Valley is a valley in Joshua Tree National Park. It has been called "the centerpiece of the park" because of the quality of its stands of Joshua trees and its geological formations. It contains the Lost Horse Mine
Lost Horse Mine is ...
in
Joshua Tree National Park
Joshua Tree National Park is an American national park in southeastern California, east of San Bernardino and Los Angeles and north of Palm Springs. It is named after the Joshua trees (''Yucca brevifolia'') native to the Mojave Desert. Original ...
. Other regions with large populations of the tree can be found northeast of
Kingman, Arizona
Kingman is a city in, and the county seat of, Mohave County, Arizona, United States. It is named after Lewis Kingman, an engineer for the Atlantic and Pacific Railroad. It is located southeast of Las Vegas, Nevada, and northwest of Arizona' ...
in Mohave County; and along U.S. 93 between the towns of Wickenburg and Wikieup, a route which has been designated the Joshua Tree Parkway of Arizona. The common name Joshua tree apparently comes from Christian iconography.
Taxonomy
The Joshua tree is also called ''izote de desierto'' (Spanish, "desert dagger"). It was first formally described in the botanical literature as ''Yucca brevifolia'' by
George Engelmann
George Engelmann, also known as Georg Engelmann, (2 February 1809 – 4 February 1884) was a German-American botanist. He was instrumental in describing the flora of the west of North America, then very poorly known to Europeans; he was particu ...
in 1871 as part of the Geological Exploration of the 100th meridian (or " Wheeler Survey").
The name "Joshua tree" is commonly said to have been given by a group of
Mormon
Mormons are a religious and cultural group related to Mormonism, the principal branch of the Latter Day Saint movement started by Joseph Smith in upstate New York during the 1820s. After Smith's death in 1844, the movement split into seve ...
settlers crossing the Mojave Desert in the mid-19th century: The tree's role in guiding them through the desert combined with its unique shape reminded them of a biblical story in which
Joshua
Joshua () or Yehoshua ( ''Yəhōšuaʿ'', Tiberian: ''Yŏhōšuaʿ,'' lit. ' Yahweh is salvation') ''Yēšūaʿ''; syr, ܝܫܘܥ ܒܪ ܢܘܢ ''Yəšūʿ bar Nōn''; el, Ἰησοῦς, ar , يُوشَعُ ٱبْنُ نُونٍ '' Yūšaʿ ...
keeps his hands reached out for an extended period of time to enable the Israelites in their conquest of Canaan (). Further, the shaggy leaves may have provided the appearance of a beard. However, no direct or contemporary attestation of this origin exists, and the name Joshua tree is not recorded until after Mormon contact; moreover, the physical appearance of the Joshua tree more closely resembles a similar story told of Moses.
Ranchers and miners who were contemporaneous with the Mormon immigrants used the trunks and branches as fencing and for fuel for ore-processing steam engines. They referred to these fallen or collapsed Joshua trees as ''tevis'' stumps.
In addition to the autonymicsubspecies ''Y. b.'' subsp. ''brevifolia'', two other subspecies have been described: ''Y. b.'' subsp. ''herbertii'' (Webber's yucca or Herbert Joshua tree) and ''Y. b.'' subsp. ''jaegeriana'' (the Jaeger Joshua tree or Jaeger's Joshua tree or pygmae yucca), though both are sometimes treated as varieties or forms. ''Y.b.'' subsp. ''jaegeriana'' has also been treated as its own species.
Growth and development
Joshua trees are fast growers for a desert species; new seedlings may grow at an average rate of per year in their first 10 years, then only about per year. The trunk consists of thousands of small
fiber
Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorpora ...
s and lacks annual growth rings, making determining the tree's age difficult. This tree has a top-heavy branch system, but also what has been described as a "deep and extensive" root system, with roots reaching down to . If it survives the rigors of the desert, it can live for hundreds of years; some specimens survive a thousand years. The tallest trees reach about . New plants can grow from
seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
, but in some populations, new stems grow from underground rhizomes that spread out around the parent tree.
The
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
leaves are dark green, linear, bayonet-shaped, 15–35 cm long, and 7–15 mm broad at the base, tapering to a sharp point; they are borne in a dense spiral arrangement at the apex of the stems. The leaf margins are white and
serrate
Serration is a saw-like appearance or a row of sharp or tooth-like projections. A serrated cutting edge has many small points of contact with the material being cut. By having less contact area than a smooth blade or other edge, the applied ...
d.
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
s typically appear from February to late April, in
panicle
A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is ...
s 30–55 cm tall and 30–38 cm broad, the individual flowers erect, 4–7 cm tall, with six creamy white to green
tepal
A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s. The tepals are
lanceolate
The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regula ...
and are fused to the middle. The fused
pistil
Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pis ...
s are 3 cm tall and the stigma cavity is surrounded by lobes. The semi-fleshy fruit that is produced is green-brown, elliptical, and contains many flat seeds. Joshua trees usually do not branch until after they bloom (though branching may also occur if the growing tip is destroyed by the yucca-boring
weevil
Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...
), and they do not bloom every year. Like most desert plants, their blooming depends on rainfall at the proper time. They also need a winter freeze before they bloom.
Once they bloom, the flowers are pollinated by the yucca moth ('' Tegeticula synthetica''), which spreads pollen while laying eggs inside the flower. The
larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Th ...
e feed on the seeds, but enough seeds remain to reproduce. The Joshua tree is also able to actively abort ovaries in which too many eggs have been produced.
Distribution and habitat
The Joshua tree is native to the southwestern United States (Arizona, California, Nevada, and Utah) and northwestern Mexico. This range mostly coincides with the geographical reach of the Mojave Desert, where it is considered one of the major
indicator species
A bioindicator is any species (an indicator species) or group of species whose function, population, or status can reveal the qualitative status of the environment. The most common indicator species are animals. For example, copepods and other sma ...
for the desert. It occurs at elevations between .
Conservation status
Joshua trees are one of the species predicted to have their range reduced and shifted by
climate change
In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
. Concern remains that they will be eliminated from Joshua Tree National Park, with ecological research suggesting a high probability that their populations will be reduced by 90% of their current range by the end of the 21st century, thus fundamentally transforming the ecosystem of the park. Wildfires, invasive grasses and poor migration patterns for the trees’ seeds are all additional factors in the species’ imperilment. As an example, more than a quarter—or more than 1.3 million Joshua trees—in of one of the densest Joshua tree populations in the world in
Mojave National Preserve
Mojave National Preserve is a United States National Preserve located in the Mojave Desert of San Bernardino County, California, USA, between Interstate 15 and Interstate 40. The preserve was established October 31, 1994, with the passage of ...
were killed in the
Dome Fire
The Dome Fire was a destructive wildfire in the Jemez Mountains in the northern region of the U.S. state of New Mexico during the 1996 fire season.
It has been described by forester Bill Armstrong as "a wakeup call that nobody woke up to", anoma ...
in August 2020. Also, concern exists about the ability of the species to migrate to favorable climates due to the extinction of the giant
Shasta ground sloth
''Nothrotheriops'' is a genus of Pleistocene ground sloth found in North America, from what is now central Mexico to the southern United States. This genus of bear-sized xenarthran was related to the much larger, and far more famous ''Megatherium ...
(''Nothrotheriops shastensis'') 13,000 years ago; ground sloth dung has been found to contain Joshua tree leaves, fruits, and seeds, suggesting that the sloths might have been key to the trees' dispersal.
Uses and cultivation
Different forms of the species are cultivated, including smaller plants native from the eastern part of the species range. These smaller plants grow 2.5 m tall and branch when about 1 m tall.
Red-shafted flicker
The northern flicker or common flicker (''Colaptes auratus'') is a medium-sized bird of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, and the Cayman Islands, and is one of the few woodpecker speci ...
s make nests in the branches, which are later used by other birds.
Cahuilla
The Cahuilla , also known as ʔívil̃uqaletem or Ivilyuqaletem, are a Native American people of the various tribes of the Cahuilla Nation, living in the inland areas of southern California.saponin glycosides.