Jewel Scarab
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The ruteline genus ''Chrysina'', or jewel scarabs (not to be confused with
jewel beetle Buprestidae is a family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy iridescent colors. Larvae of this family are known as flatheaded borers. The family is among the largest of the beetles, with some ...
s, which are a different family), is a large genus of brightly colored, often metallic
iridescent Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear to gradually change color as the angle of view or the angle of illumination changes. Examples of iridescence include soap bubbles, feathers, butterfl ...
species, ranging from the southwestern edge of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
as far south as
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
and
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
. The genus includes all the species formerly known as ''Plusiotis''.Hawks, D.C. 2001. Taxonomic and nomenclatural changes in ''Chrysina'' and a synonymic checklist of species (Scarabaeidae: Rutelinae). Occ. Papers Consort. Coleopt. 4: 1-8 They are typically between in length, and are nocturnal in habits, coming readily to lights. The larvae live in rotting logs, while the adults commonly feed on foliage; they tend to be found in
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
,
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
, or
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
-
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
forests, most commonly between elevation. They are most diverse in countries such as
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
, where as many as 15 species can be found in a single location, but there are only 4 species which occur in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. These beetles are very popular among collectors; many species are polymorphic, with rare color variants that can command high prices. One such specimen was featured on the cover of National Geographic Magazine. The majority of species are bright green, but metallic silver and gold are also common colors, and may be combined with green as in the common ''
Chrysina gloriosa ''Chrysina gloriosa'' is a species of scarab whose common names are glorious beetle and glorious scarab. The adults are long and are bright green with silver stripes on the elytra. These iridescent stripes on the cuticle of the elytra are a res ...
'' from the Madrean sky islands region. Particularly attractive specimens were reported to sell for as much as $500 in 2007. The genus name is from Greek χρύσινος (''chrysinos''), "gold-coloured".


Species

*'' Chrysina adelaida'' (Hope, 1841) *'' Chrysina adolphi'' Chevrolat, 1859 *'' Chrysina alexae'' Monzón, 2017 *'' Chrysina alfredolaui'' (Hawks, 1995) *'' Chrysina alphabarrerai'' (Morón, 1981) *'' Chrysina amalia'' (Burmeister, 1844) *'' Chrysina antonkozlovi'' Monzón, 2017 *'' Chrysina arellanoi'' Monzón, 2012 *'' Chrysina argenteola'' (H. Bates, 1888) *'' Chrysina aurigans'' (Rothschild & Jordan, 1894) *'' Chrysina aurilisternum'' Perez-Flores, Villagomez & Galindo, 2016 *'' Chrysina auripes'' Gray, 1832 *'' Chrysina aurofoveata'' (Morón, 1981) *'' Chrysina auropunctata'' (Ohaus, 1913) *'' Chrysina aurora'' (Boucard, 1875) *'' Chrysina badeni'' (Boucard, 1878) *'' Chrysina baileyana'' Monzón, 2010 *'' Chrysina batesi'' (Boucard, 1875) *'' Chrysina beckeri'' H. Bates, 1889 *'' Chrysina benesi'' Pokorný & Curoe, 2012 *'' Chrysina beraudi'' (Warner, Hawks & Bruyea, 1992) *'' Chrysina beyeri'' (Skinner, 1905) *'' Chrysina blackalleri'' Monzón & García, 2011 *'' Chrysina boucardi'' (Sallé, 1878) *'' Chrysina brevis'' (Rothschild & Jordan, 1894) *'' Chrysina bruyeai'' (Hawks, 1999) *'' Chrysina cavei'' Hawks & Bruyea, 1999 *'' Chrysina centralis'' (Morón, 1990) *'' Chrysina chalcothea'' (H. Bates, 1888) *'' Chrysina chimalapensis'' Mora-Aguilar, Curoe, Delgado & Ramírez-Ponce 2018 *'' Chrysina chloreis'' (H. Bates, 1888) *'' Chrysina chrysargyrea'' (Sallé, 1874) *'' Chrysina chrysopedila'' (H. Bates, 1888) *'' Chrysina citlaltepetlamayatli'' (Blackaller-Bages & Delgado, 1994) *'' Chrysina clavellina'' Monzón, Blackaller, & Hawks, 2020 *'' Chrysina clypealis'' (Rothschild & Jordan, 1894) *'' Chrysina colima'' (Morón, 1992) *'' Chrysina confusa'' (Ohaus, 1913) *'' Chrysina costata'' (Blanchard, 1850) *'' Chrysina crassimargo'' (Rothschild & Jordan, 1894) *'' Chrysina cunninghami'' (Curoe, 1999) *'' Chrysina cupreomarginata'' (F. Bates, 1904) *'' Chrysina curoei'' (Warner, LeBlanc, Hawks & Bruyea, 1992) *'' Chrysina cusuquensis'' (Curoe, 1994) *'' Chrysina dianae'' (Ratcliffe & Taylor, 1992) *'' Chrysina difficilis'' (Morón, 1990) *'' Chrysina diversa'' (Ohaus, 1912) *'' Chrysina donthomasi'' Monzón & García, 2011 *'' Chrysina dzidorhum'' (Arnaud, 1994) *'' Chrysina ericsmithi'' (Monzón & Cano, 1999) *'' Chrysina erubescens'' H. Bates, 1889 *'' Chrysina expansa'' (Ohaus, 1913) *'' Chrysina eyai'' Curoe, 2012 *'' Chrysina falcifera'' Hawks, 2017 *'' Chrysina flohri'' (Ohaus, 1905) *'' Chrysina gaitalica'' Curoe, 2012 *'' Chrysina galbina'' Hawks, 2017 *'' Chrysina giesberti'' Monzón, 2010 *''
Chrysina gloriosa ''Chrysina gloriosa'' is a species of scarab whose common names are glorious beetle and glorious scarab. The adults are long and are bright green with silver stripes on the elytra. These iridescent stripes on the cuticle of the elytra are a res ...
'' (Leconte, 1854) *'' Chrysina gorda'' Delgado, 2003 *'' Chrysina guatemalensis'' (Monzón, Cano & Bailey, 1999) *'' Chrysina guaymi'' (Curoe, 2001) *'' Chrysina halffteri'' (Morón, 1990) *'' Chrysina hawksi'' Monzón, 2010 *'' Chrysina howdenorum'' (Morón, 1990) *'' Chrysina intermedia'' (Ohaus, 1913) *'' Chrysina juxtaprasina'' Hawks, 2017 *'' Chrysina kalinini'' Zubov & Ivshin, 2019 *'' Chrysina karschi'' (Nonfried, 1891) *'' Chrysina lacordairei'' (Boucard, 1875) *'' Chrysina laniventris'' (Sturm, 1843) *'' Chrysina lecontei'' (Horn, 1882) *''
Chrysina limbata ''Chrysina limbata'' is a species of scarab beetle found in the tropical rainforests of Central America, including Costa Rica, and Mexico. It is in the genus '' Chrysina'', in the subfamily Rutelinae (shining leaf chafers). It is notable for it ...
'' (Rothschild & Jordan, 1894) *'' Chrysina luteomarginata'' (Ohaus, 1913) *'' Chrysina macropus'' (Francillon, 1795) *'' Chrysina magnistriata'' (Morón, 1990) *'' Chrysina maishei'' Monzón, 2017 *'' Chrysina marginata'' (Waterhouse, 1871) *'' Chrysina miguelangeli'' Nogueira & Curoe, 2012 *'' Chrysina misteca'' (Morón, 1990) *'' Chrysina modesta'' (Sturm, 1843) *'' Chrysina moroni'' (Curoe & Beraud, 1994) *'' Chrysina nogueirai'' (Morón, 1992) *'' Chrysina ofidiodontophallica'' Curoe, 2011 *'' Chrysina optima'' (H. Bates, 1888) *'' Chrysina oreicola'' (Morón, 1992) *'' Chrysina orizabae'' (H. Bates, 1889) *'' Chrysina pastori'' (Curoe, 1994) *'' Chrysina paulseni'' Hawks, 2017 *'' Chrysina pehlkei'' (Ohaus, 1930) *'' Chrysina peruviana'' Kirby, 1828 *'' Chrysina plusiotina'' (Ohaus, 1912) *'' Chrysina porioni'' Monzón & Hawks, 2020 *'' Chrysina prasina'' (Boucard, 1878) *'' Chrysina pricei'' Hawks, 2020 *'' Chrysina prototelica'' (Morón & Howden, 1992) *'' Chrysina psittacina'' (Sturm, 1843) *'' Chrysina purpurata'' (Morón, 1990) *'' Chrysina purulhensis'' (Monzón & Warner, 1993) *'' Chrysina quetzalcoatli'' (Morón, 1990) *'' Chrysina quiche'' (Morón, 1990) *'' Chrysina ratcliffei'' (Morón, 1990) *''
Chrysina resplendens ''Chrysina resplendens'' is a scarab beetle found in Costa Rica, Panama, El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is b ...
'' (Boucard, 1875) *'' Chrysina robackeri'' Hawks, 2020 *'' Chrysina rodriguezi'' (Boucard, 1878) *'' Chrysina sagacita'' Hawks, 2017 *'' Chrysina sallaei'' (Boucard, 1875) *'' Chrysina schusteri'' (Monzon, Cano & Bailey, 1999) *'' Chrysina sirenicola'' (Solís & Morón, 1995) *'' Chrysina spectabilis'' (Ratcliffe & Jameson, 1992) *'' Chrysina strasseni'' (Ohaus, 1924) *'' Chrysina tapantina'' (Morón, 1992) *'' Chrysina taylori'' (Morón, 1990) *'' Chrysina tecunumani'' (Cano & Morón, 1995) *'' Chrysina terroni'' (Morón, 1990) *'' Chrysina transvolcanica'' (Morón & Nogueira, 2016) *'' Chrysina tricolor'' (Ohaus, 1922) *'' Chrysina triumphalis'' Morón, 1990 *'' Chrysina tuerckheimi'' (Ohaus, 1913) *'' Chrysina valentini'' Zubov & Ivshin, 2019 *'' Chrysina veraguana'' (Ohaus, 1922) *'' Chrysina victorina'' (Hope, 1841) *'' Chrysina wolfi'' (Ohaus, 1912) *'' Chrysina woodi'' (Horn, 1884) *'' Chrysina woodruffi'' Monzón, 2017 *'' Chrysina xalixteca'' (Morón, 1992) *'' Chrysina zapoteca'' (Morón, 1990)


Metallic color mechanism

Research has shown that the elytra of ''Chrysina'' are composed of about 70
chitin Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chit ...
ous layers of exoskeleton. The different reflective indices and spacing of the layers cause light of different wavelengths to be selectively refracted through them and by them in different phases, leading to a metallic appearance, and also leads to different specimens having different colors.


References


External links


Generic Guide to New World Scarab Beetles
{{Taxonbar, from=Q11979610 Rutelinae