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''Jeffersonia'', also known as twinleaf or rheumatism root, is a small
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s in the family
Berberidaceae The Berberidaceae are a family of 18 genera of flowering plants commonly called the barberry family. This family is in the order Ranunculales. The family contains about 700 known species, of which the majority are in ''Berberis''. The species i ...
. They are uncommon spring wildflowers and grow in
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
soils of rich
deciduous forests In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
. ''Jeffersonia'' was named for United States President Thomas Jefferson by his contemporary Benjamin Smith Barton. This genus was formerly grouped in genus '' Podophyllum''. Twinleaf is protected by state laws as a threatened or endangered plant in Georgia, Iowa,
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
, and New Jersey.


Description

''Jeffersonia diphylla'' has leaves and flowers that are smooth and emerge directly from the rhizome. The leaves are divided into two leaflets which are half-ovate in shape with entire or shallowly toothed margins. It has showy white flowers with four sepals, eight petals, eight stamens, and a singly pistil; the flower resembles bloodroot flowers. The short-lived flower appears in April or May before the forest canopy appears (see
spring ephemeral An ephemeral plant is one marked by short life cycles. The word ephemeral means transitory or quickly fading. In regard to plants, it refers to several distinct growth strategies. The first, spring ephemeral, refers to perennial plants that emerge ...
). The fruit is a green pear-shaped capsule with a hinged top. The characteristic leaves are large and nearly divided in half, giving rise to its common name, twinleaf. It rarely grows taller than . As with other deciduous forest plants, the seeds are dispersed by ants, a process known as myrmecochory.


Species

;accepted speciesThe Plant List, search for ''Jeffersonia''
/ref> *''Jeffersonia diphylla'' (L.) Pers. – Eastern North America especially Great Lakes region, Ohio Valley, and Appalachians ;unresolved names *''Jeffersonia dubia'' (Maxim.) Benth. & Hook. f. ex Baker & Moore – China, Korea, Russia (called ''
Plagiorhegma dubium ''Plagiorhegma'' is a monotypic genus of flowering plants belonging to the family Berberidaceae. The only species is ''Plagiorhegma dubium'', the Asian twinleaf. Its native range is Russian Far East The Russian Far East (russian: ДальР...
'' in Flora of China) *''Jeffersonia lobata'' Nutt. *''Jeffersonia odorata'' Raf. ;species in homonymic genus In 1800, Brickell used the name ''Jeffersonia'' to refer to some plants in the Loganiaceae, thus creating an illegitimate homonym. Species names coined using this illegitimate use of the name: *''Jeffersonia sempervirens'' Brickell, now called '' Gelsemium sempervirens'' (L.) J.St.-Hil


Distribution and habitat

''Jeffersonia diphylla'' is native to Eastern North America where it is found from Minnesota east to New York and south to Tennessee. In Minnesota it grows in the lower southeastern section of the state in mesic deciduous forests and most commonly in ravines or on talus slopes. In Minnesota it has a preference for growing on calcareous soils of north and east-facing slopes and is listed as a threatened species. In Minnesota it is found in association with other uncommon woodland plants such as ''Hydrastis canadensis'' (
golden seal ''For the magazine from West Virginia see Goldenseal (magazine)'' Goldenseal (''Hydrastis canadensis''), also called orangeroot or yellow puccoon, is a perennial herb in the buttercup family Ranunculaceae, native to North America. It may be dis ...
) and ''Dicentra canadensis'' ( squirrel-corn).


Uses

''Jeffersonia'' has had a variety of medical uses. One is hinted at by an archaic common name of ''Jeffersonia diphylla'', Rheumatism root. The roots of both species contain berberine, a known anti-tumor alkaloid. The plant is therefore considered poisonous.


America

Native Americans use ''Jeffersonia diphylla'' for a variety of medicines. The Cherokee use an infusion of this plant for treating dropsy, as well as gravel and urinary tract problems, and as a poultice for sores and inflammation. The Iroquois used a decoction of the plant to treat
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to be ...
and diarrhea. The whole plant was used in early American medicine as an antispasmodic, diuretic, emetic, expectorant and general tonic. The "root" was once also used as an emetic in large doses, and as an expectorant in small doses.Plants for a Future Database: ''J. diphylla''
/ref> Modern medicine does not use this plant.


References

{{Taxonbar, from=Q2562522 Berberidaceae Berberidaceae genera Monotypic Ranunculales genera Thomas Jefferson Taxa named by Benjamin Smith Barton