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The Javan leopard (''Panthera pardus melas'') is a
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
subspecies confined to the
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
n island of
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
. It has been listed as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
since 2021. The population is estimated at 188–571 mature individuals in 22 fragmented subpopulations and a declining population trend. The total remaining habitat is estimated at only .


Characteristics

The Javan leopard was initially described as a
black panther A black panther is the melanistic colour variant of the leopard (''Panthera pardus'') and the jaguar (''Panthera onca''). Black panthers of both species have excess black pigments, but their typical rosettes are also present. They have been d ...
with dark black spots and silver-grey eyes. It has either a normal spotted coat with rosettes, or a
recessive In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and t ...
phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
resulting in a black coat.


Distribution and habitat

The Javan leopard is confined to the Indonesian island of Java. It is known to inhabit
Gunung Halimun National Park Mount Halimun Salak National Park is a 400 km2 conservation area in the Indonesian province of West Java on the island of Java. Established in 1992, the park comprises two mountains, Mount Salak and Mount Halimun with an 11-kilometer forest c ...
,
Ujung Kulon National Park Ujung Kulon National Park is at the westernmost tip of Java, in Banten province of Indonesia. It includes the volcanic island group of Krakatoa in Lampung province, and other islands including Panaitan, as well as smaller offshore islets such as ...
,
Gunung Gede Pangrango National Park Mount Gede Pangrango National Park is a national park in West Java, Indonesia. The park is centred on two volcanoes—Mount Gede and Mount Pangrango—and is 150 km² in area. It evolved from already existing conservation areas, such as Ci ...
, Ceremai National Park,
Merbabu Mount Merbabu ( id, Gunung Merbabu) is a dormant stratovolcano in Central Java province on the Indonesian island of Java. The name ''Merbabu'' could be loosely translated as 'Mountain of Ash' from the Javanese combined words; ''Meru'' means " ...
National Park, Merapi National Park,
Bromo Tengger Semeru National Park Bromo Tengger Semeru National Park ( id, Taman Nasional Bromo Tengger Semeru; abbreviated as ) is a national park located in East Java, Indonesia, to the east of Malang and Lumajang, to the south of Pasuruan and Probolinggo, and to the south ...
,
Meru Betiri National Park Meru Betiri National Park is a national park in the province of East Java, Indonesia, extending over an area of 580 km2 of which a small part is marine (8.45 km2). The beaches of the park provide nesting grounds for endangered turtle spe ...
, Ijen Mountain,
Baluran National Park Baluran National Park is located in Situbondo Regency, East Java, Indonesia. It has a relatively dry climate and mainly consists of savanna (40%), as well as lowland forests, mangrove forests and hills, with Mount Baluran (1,247m) as its highest ...
, and
Alas Purwo National Park Alas Purwo National Park is situated on Blambangan Peninsula in the Banyuwangi Regency, at the southeastern tip of East Java province. The park is famous for its wild Banteng and surfing location at Grajagan Bay. The park's name means ''first ...
. It inhabits altitudes from sea level to ranging from dense
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equatori ...
to dry
deciduous forest In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
s. Outside protected areas, it was recorded in
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
, mixed
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to ...
and production forest between 2008 and 2014. In the 1990s, it survived in the
seral A seral community is an intermediate stage found in ecological succession in an ecosystem advancing towards its climax community. In many cases more than one seral stage evolves until climax conditions are attained. A prisere is a collection of se ...
stages of successional vegetation patterns, which made it less susceptible to humans' disruptive activities than many other mammals. From 2001 to 2004, monitoring research has been conducted in a area of Gunung Halimun National Park using
camera trap A camera trap is a camera that is automatically triggered by a change in some activity in its vicinity, like presence of an animal or a human being. It is typically equipped with a motion sensor – usually a passive infrared (PIR) sensor ...
s and radio tracking. Seven leopards were identified in the study area. The total population was estimated at 42 to 58 individuals. The home range of an adult female averaged .Harahap, S., Sakaguchi, H. (2005). ''Ecological research and conservation of the Javan Leopard ''Panthera pardus melas'' in Gunung Halimun National Park, West Java, Indonesia''. In
The wild cats: ecological diversity and conservation strategy
The 21st Century Center of Excellence Program International Symposium. Okinawa, Japan.


Ecology and behavior

The Javan leopard's prey comprises
barking deer Barking may refer to: Places * Barking, London, a town in East London, England ** London Borough of Barking and Dagenham, a local government district covering the town of Barking ** Municipal Borough of Barking, a historical local government dist ...
,
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
,
Java mouse-deer The Java mouse-deer (''Tragulus javanicus'') is a species of even-toed ungulate in the family Tragulidae. When it reaches maturity it is about the size of a rabbit, making it the smallest living ungulate. It is found in forests in Java and perhap ...
, and primates such as
crab-eating macaque The crab-eating macaque (''Macaca fascicularis''), also known as the long-tailed macaque and referred to as the cynomolgus monkey in laboratories, is a cercopithecine primate native to Southeast Asia. A species of macaque, the crab-eating macaqu ...
,
silvery lutung The silvery lutung (''Trachypithecus cristatus''), also known as the silvered leaf monkey or the silvery langur, is an Old World monkey. It is arboreal, living in coastal, mangrove, and riverine forests in Peninsular Malaysia, Sumatra, and Borne ...
, and
Javan gibbon The silvery gibbon (''Hylobates moloch''), also known as the Javan gibbon, is a primate in the gibbon family Hylobatidae. It is endemic to the Indonesian island of Java, where it inhabits undisturbed rainforests up to an altitude of . It is liste ...
. Javan leopards also look for food in close by villages and have been known to prey on domestic dogs, chickens, and goats. Two leopards were radio collared in the Gunung Halimun National Park. Their daily activity pattern showed peaks in the early mornings between 6:00 and 9:00, and late afternoons between 15:00 and 18:00.


Threats

The Javan leopard is threatened by loss of habitat, prey base depletion, and
poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
due to human population growth and
agricultural expansion Agricultural expansion describes the growth of agricultural land (arable land, pastures, etc.) especially in the 20th and 21st centuries. The agricultural expansion is often explained as a direct consequence of the global increase in food and ene ...
. Conflict between local people and leopards is also considered to be a main threat to the Javan leopard. Java has lost more than 90% of its natural vegetation and is one of the most densely populated islands in the world. Primary forests remain only in the mountainous regions at elevations above .


Conservation

''Panthera pardus melas'' is listed in the
CITES Appendix I CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
. Efforts are being made to restore the Javan leopard population and prevent its extinction. Hunting laws are strictly enforced. In 2005, Gunung Halimun National Park was enlarged to three times its original size for the protection of the Javan leopard, the
silvery gibbon The silvery gibbon (''Hylobates moloch''), also known as the Javan gibbon, is a primate in the gibbon family Hylobatidae. It is endemic to the Indonesian island of Java, where it inhabits undisturbed rainforests up to an altitude of . It is liste ...
(''Hylobates moloch''), and the
Javan hawk-eagle The Javan hawk-eagle (''Nisaetus bartelsi'') is a medium-sized, dark brown raptor in the family Accipitridae. It is the national bird of Indonesia, where it is commonly referred to as Garuda, from the bird-like creatures in Hindu and Buddhist myt ...
(''Nisaetus bartelsi''). To address the issue of Java's overpopulation and encroachment on habitat of protected species, the Indonesian government has formed a nationwide family planning program. This program makes contraceptive devices like condoms and several different forms of birth control pills more readily available to the public.


In captivity

In 1997, 14 Javan leopards were kept in European zoos. The Javan leopard is not specifically managed in
captive breeding Captive breeding, also known as captive propagation, is the process of plants or animals in controlled environments, such as wildlife reserves, zoos, botanic gardens, and other conservation facilities. It is sometimes employed to help species that ...
programs in Europe and America. As of 2007, the
Taman Safari Taman Safari Indonesia or simply Taman Safari are animal theme parks in Cisarua, Prigen, and Bali. Being part of the same organization, they are known as Taman Safari I, II and III. The most popular is Taman Safari I. Taman Safari I Taman ...
Zoo in
Bogor Bogor ( su, , nl, Buitenzorg) is a city in the West Java province, Indonesia. Located around south of the national capital of Jakarta, Bogor is the 6th largest city in the Jakarta metropolitan area and the 14th overall nationwide.Ragunan Ragunan is a '' kelurahan'' (Administrative Village) of Pasar Minggu Subdistrict, South Jakarta South Jakarta ( id, Jakarta Selatan; bew, Jakarte Beludik ), colloquially known as ''Jaksel'', is one of the five administrative cities which form ...
,
Taman Safari Taman Safari Indonesia or simply Taman Safari are animal theme parks in Cisarua, Prigen, and Bali. Being part of the same organization, they are known as Taman Safari I, II and III. The most popular is Taman Safari I. Taman Safari I Taman ...
and Surabaya Zoo also keep Javan leopards. As of December 2011, two male and one female Javan leopard were kept in
Tierpark Berlin The Tierpark Berlin is one of two zoos located in Berlin, Germany. It was founded in 1955 and is located in Friedrichsfelde on the former grounds of Friedrichsfelde Palace, which is situated within the zoo. , the zoo houses 7,250 animals from ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
; and one male and one female in the
Jakarta Jakarta (; , bew, Jakarte), officially the Special Capital Region of Jakarta ( id, Daerah Khusus Ibukota Jakarta) is the capital and largest city of Indonesia. Lying on the northwest coast of Java, the world's most populous island, Jakarta ...
Zoo. In 2013, one male Javan leopard was transferred from Tierpark Berlin to the
Prague Zoo Prague Zoological Garden (Czech: ''Zoologická zahrada hl. m. Prahy'') is a zoo in Prague, Czech Republic. It was opened in 1931 with the goal to "advance the study of zoology, protect wildlife, and educate the public" in the district of Troja in ...
.


Evolution

Morphological research indicates that the Javan leopard is craniometrically distinct from other Asian leopard subspecies, and is a distinct
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
that split off from other Asian leopard subspecies in the
Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. The ...
about 800,000 years ago. In the Middle Pleistocene, it may have migrated to Java from South Asia across a land bridge that bypassed Sumatra and Borneo.


See also

* Leopard subspecies *
Chinese leopard The term Chinese leopard refers to any of the following three leopard (''Panthera pardus'') subspecies occurring in China: *Indian leopard (''P. p. fusca'') in southern Tibet, where leopards have been recorded in Qomolangma National Nature Preserve. ...
*
Zanzibar leopard The Zanzibar leopard is an African leopard (''Panthera pardus pardus'') population on Unguja Island in the Zanzibar archipelago, Tanzania, that is considered extirpated due to persecution by local hunters and loss of habitat. It was the island's ...


References


External links

* Species portrai
''Panthera pardus'' in Asia
and short portrai
''P. pardus melas''
IUCN/SSC Cat Specialist Group
''Critically Endangered Javan Leopard Captured on Camera Traps in Ujung Kulon, Indonesia''
{{Taxonbar, from=Q841950 Leopards Critically endangered animals Critically endangered biota of Asia Endemic fauna of Indonesia Fauna of Java Mammals of Indonesia Taxa named by Georges Cuvier Apex predators