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The Indian leopard (''Panthera pardus fusca'') is a
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, ...
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
widely distributed on the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, In ...
. The
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
''Panthera pardus'' is listed as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
because populations have declined following
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and fragmentation,
poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
for the illegal trade of skins and body parts, and persecution due to conflict situations. The Indian leopard is one of the
big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. Despite enormous differences in size, various cat species are quite similar ...
s occurring on the Indian subcontinent, along with the
Asiatic lion The Asiatic lion is a population of '' Panthera leo leo'' that today survives in the wild only in India. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of Gujar ...
,
Bengal tiger The Bengal tiger is a population of the '' Panthera tigris tigris'' subspecies. It ranks among the biggest wild cats alive today. It is considered to belong to the world's charismatic megafauna. The tiger is estimated to have been present i ...
,
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...
and
clouded leopard The clouded leopard (''Neofelis nebulosa''), also called the mainland clouded leopard, is a wild cat inhabiting dense forests from the foothills of the Himalayas through mainland Southeast Asia into South China. In the early 19th century, a ...
. In 2014, a national census of leopards around tiger habitats was carried out in India except the northeast. 7,910 individuals were estimated in surveyed areas and a national total of 12,000–14,000 speculated.


Taxonomy

''Felis fusca'' was the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
proposed by Friedrich Albrecht Anton Meyer in 1794 who described a black leopard from
Bengal Bengal ( ; bn, বাংলা/বঙ্গ, translit=Bānglā/Bôngô, ) is a geopolitical, cultural and historical region in South Asia, specifically in the eastern part of the Indian subcontinent at the apex of the Bay of Bengal, predom ...
that was on display at the
Tower of London The Tower of London, officially His Majesty's Royal Palace and Fortress of the Tower of London, is a historic castle on the north bank of the River Thames in central London. It lies within the London Borough of Tower Hamlets, which is sep ...
. ''Leopardus perniger'' proposed by
Brian Houghton Hodgson Brian Houghton Hodgson (1 February 1800 or more likely 1801 – 23 May 1894) was a pioneer naturalist and ethnologist working in India and Nepal where he was a British Resident. He described numerous species of birds and mammals from the Hima ...
in 1863 were five leopard skins from
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
, out of which three were
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white ha ...
. He mentioned
Sikkim Sikkim (; ) is a state in Northeastern India. It borders the Tibet Autonomous Region of China in the north and northeast, Bhutan in the east, Province No. 1 of Nepal in the west and West Bengal in the south. Sikkim is also close to the Sil ...
and Nepal as habitat. ''Panthera pardus millardi'' proposed by
Reginald Innes Pocock Reginald Innes Pocock Fellow of the Royal Society, F.R.S. (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock (historian), Nicholas Pocock and Edith Prichard. He ...
in 1930 was a single leopard skin and skull from
Kashmir Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompas ...
. It differed from typical ''P. p. fusca'' skins by longer hair and a more greyish colour. Since leopard populations in Nepal, Sikkim and Kashmir are not geographically isolated from leopard populations in the Indian subcontinent, they were subsumed to ''P. p. fusca'' in 1996.


Characteristics

The Indian leopard has strong legs and a long well-formed tail, broad muzzle, short ears and small, yellowish grey eyes, light grey ocular bulbs. Its coat is spotted and rosetted on a pale yellow to yellowish-brown or golden background, except for the
melanistic The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair. Pseudomelanism, also called abundism, is another variant of pi ...
forms; the spots fade toward the white underbelly and the insides and lower parts of the legs. Rosettes are most prominent on the back, flanks and hindquarters. The pattern of the rosettes is unique to each individual. Juveniles have woolly fur, and appear dark due to the densely arranged spots. The white-tipped tail is long, white underneath, and displays rosettes, which form incomplete bands toward the end. The rosettes are larger in other leopard subspecies in Asia. Fur colour tends to be more pale and cream in arid habitats, more gray in colder climates, and of a darker golden hue in
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
habitats. The clouded leopard can be told apart by its diffuse "clouds" of spots compared to the smaller and distinct rosettes of the leopard, longer legs and thinner tail.


Skull

The largest skull recorded for an Indian leopard belonged to a large black panther in the area of
Ootacamund Ooty (), officially known as Udhagamandalam (also known as Ootacamund (); abbreviated as Udhagai), is a city and a municipality in the Nilgiris district of the South Indian state of Tamil Nadu. It is located north west of Coimbatore and ...
, which was recorded in 1920. The panther was said to have bigger forelimbs and forequarters than hind-limbs and hind-quarters, and a skull and claws about as large as those of a tigress. The skull measured in basal length, and in breadth, and weighed . To compare, the skull of a
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali ...
n panther measured in basal length, and in breadth, and weighed .


Size

Male Indian leopards grow to between and in body size with a to long tail and weigh between . Females are smaller, growing to between and in body size with a to long tail, and weigh between .
Sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, males are larger and heavier than females. The largest wild individual appears to have been a male man-eater that was shot in the Dhadhol area of Bilaspur district, Himachal Pradesh in 2016. It reportedly measured from head to tail, at the shoulder, and weighed .


Distribution and habitat

The Indian leopard is distributed in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
and parts of
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
. Bangladesh has no viable leopard population but there are occasional sightings in the forests of
Sylhet Sylhet ( bn, সিলেট) is a metropolitan city in northeastern Bangladesh. It is the administrative seat of the Sylhet Division. Located on the north bank of the Surma River at the eastern tip of Bengal, Sylhet has a subtropical climate ...
,
Chittagong Hill Tracts The Chittagong Hill Tracts ( bn, পার্বত্য চট্টগ্রাম, Parbotto Chottogram), often shortened to simply the Hill Tracts and abbreviated to CHT, are group of districts within the Chittagong Division in southeast ...
and
Cox's Bazar Cox's Bazar (; bn, কক্সবাজার, Kôksbajar; ) is a city, fishing port, tourism centre, and district headquarters in Southeastern Bangladesh. It is located south of the city of Chittagong. Cox's Bazar is also known by the n ...
. It inhabits
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equator ...
s, dry
deciduous forest In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals ...
s,
temperate forest A temperate forest is a forest found between the tropical and boreal regions, located in the temperate zone. It is the second largest biome on our planet, covering 25% of the world's forest area, only behind the boreal forest, which covers abou ...
s and northern
coniferous forest Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ...
s but does not occur in the
mangrove forest Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fre ...
s of the
Sundarbans Sundarbans (pronounced ) is a mangrove area in the delta formed by the confluence of the Padma, Brahmaputra and Meghna Rivers in the Bay of Bengal. It spans the area from the Baleswar River in Bangladesh's division of Khulna to the Hooghly R ...
. It is thought that the
Indus River The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmi ...
in the west and the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
in the north form topographical barriers to the dispersal of this subspecies. In the east, the Ganges Delta and the lower course of the
Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. I ...
are thought to form natural barriers to the range of the Indochinese leopard. In southern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
, it was recorded in Qomolangma National Nature Preserve. In Pakistan, it inhabits
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
n forests and mountainous regions. In the 1970s, it was still recorded in the Kirthar Mountains, northeastern
Baluchistan Balochistan ( ; bal, بلۏچستان; also romanised as Baluchistan and Baluchestan) is a historical region in Western Asia, Western and South Asia, located in the Iranian plateau's far southeast and bordering the Indian Plate and the Arabian S ...
and
Murree Hills Murree ( Punjabi, Urdu: مری) is a mountain resort city, located in the Galyat region of the Pir Panjal Range, within the Muree District of Punjab, Pakistan. It forms the outskirts of the Islamabad-Rawalpindi metropolitan area, and is about ...
. Since the turn of the century, leopards were recorded in and around Machiara National Park, Pir Lasora National Park, and Ayubia National Park. Leopards are occasionally spotted in the
Margalla Hills The Margalla Hills () are a hill range within the Margalla Hills National Park on the northern edge of Islamabad Capital Territory, Pakistan, just south of Haripur District, Khyber Pakhtunkhwa. They are part of the Himalayan foothills. Th ...
in winter and are observed preying on monkeys in the woods as well as on local livestock. In April 2020, photos of three leopard families were taken by camera traps in the Margalla Hills. In Nepal's
Kanchenjunga Conservation Area Kanchenjunga Conservation Area is a protected area in the Himalayas of eastern Nepal that was established in 1997. It covers in the Taplejung District and comprises two peaks of Kanchenjunga. In the north it adjoins the Qomolangma National Nature ...
, a melanistic leopard was photographed at an elevation of by a camera trap in May 2012.


Population in India

In 2015, 7,910 leopards were estimated to live in and around tiger habitat in India; about 12,000 to 14,000 leopards were speculated to live in the entire country. The following table gives the major leopard populations in the Indian states. As of 2020, the leopard population within forested habitats in India's tiger range landscapes was estimated at 12,172 to 13,535 individuals. Surveyed landscapes included elevations below in the
Shivalik Hills The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Ind ...
and
Gangetic plains The Indo-Gangetic Plain, also known as the North Indian River Plain, is a fertile plain encompassing northern regions of the Indian subcontinent, including most of northern and eastern India, around half of Pakistan, virtually all of Ba ...
,
Central India Central India is a loosely defined geographical region of India. There is no clear official definition and various ones may be used. One common definition consists of the states of Chhattisgarh and Madhya Pradesh, which are included in al ...
and
Eastern Ghats The Eastern Ghats are a discontinuous range of mountains along India's eastern coast. The Eastern Ghats pass through Odisha, Andhra Pradesh to Tamil Nadu in the south passing some parts of Karnataka as well as Telangana. They are eroded and ...
, Western Ghats, as well as the
Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. I ...
basin and hills in
Northeast India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...
.


Behaviour and ecology

The leopard is elusive, solitary, and largely
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
. It is known for its ability in climbing, and has been observed resting on tree branches during the day, dragging its kills up trees and hanging them there, and descending from trees headfirst. It is a powerful swimmer, although is not as disposed to swimming as the tiger. It is very agile, and can run at over , leap over horizontally, and jump up to vertically. It produces a number of vocalizations, including grunts, roars, growls, meows, and purrs. In Nepal's Bardia National Park, home ranges of male leopards comprised about , and of females about ; female home ranges decreased to when they had young cubs. In Gir National Park, the home range of a male radio-collared leopard was estimated at . It killed prey once in 3.7 days. The leopard is a versatile, opportunistic hunter, and has a very broad diet. It is able to take large prey due to its massive skull and powerful jaw muscles. In Sariska Tiger Reserve, the dietary spectrum of the Indian leopard includes axis deer,
sambar deer The sambar (''Rusa unicolor'') is a large deer native to the Indian subcontinent and Southeast Asia that is listed as a vulnerable species on the IUCN Red List since 2008. Populations have declined substantially due to severe hunting, local ins ...
,
nilgai The nilgai (''Boselaphus tragocamelus'') (, literally meaning "blue cow") is the largest Asian antelope and is ubiquitous across the northern Indian subcontinent. It is the sole member of the genus ''Boselaphus'' and was described by Peter Si ...
,
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species i ...
,
common langur Gray langurs, also called Hanuman langurs and Hanuman monkeys, are Old World monkeys native to the Indian subcontinent constituting the genus ''Semnopithecus''. Traditionally only one species ''Semnopithecus entellus'' was recognized, but since a ...
,
Indian hare The Indian hare (''Lepus nigricollis''), also known as the black-naped hare, is a common species of hare native to the Indian subcontinent, and Java. Introductions It has been introduced to Madagascar, Comoro Islands, Andaman Islands, Irian Jay ...
and
peafowl Peafowl is a common name for three bird species in the genera '' Pavo'' and '' Afropavo'' within the tribe Pavonini of the family Phasianidae, the pheasants and their allies. Male peafowl are referred to as peacocks, and female peafowl are ref ...
. In
Periyar Tiger Reserve Periyar National Park and Wildlife Sanctuary (PNP) is a protected area located in the districts of Idukki and Pathanamthitta in Kerala, India. It is notable as an elephant reserve and a tiger reserve. The protected area encompasses , of whi ...
,
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
s make up a large proportion of its diet.


Reproduction

Depending on the region, the leopard mates all year round. The
estrous cycle The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestro ...
lasts about 46 days and the female usually is in heat for 6–7 days. Gestation lasts for 90 to 105 days. Cubs are usually born in a litter of 2–4 cubs. Mortality of cubs is estimated at 41–50% during the first year. Females give birth in a cave, crevice among boulders, hollow tree, or thicket to make a den. Cubs are born with closed eyes, which open four to nine days after birth. The fur of the young tends to be longer and thicker than that of adults. Their pelage is also more gray in colour with less defined spots. Around three months of age, the young begin to follow the mother on hunts. At one year of age, leopard young can probably fend for themselves, but remain with the mother for 18–24 months. The average typical life span of a leopard is between 12 and 17 years.


Sympatric carnivores

Indian leopards are not common in habitats where
tiger The tiger (''Panthera tigris'') is the largest living Felidae, cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily pr ...
density is high, and are wedged between prime tiger habitat on the one side, and cultivated village land on the other. Where the tiger population is high or increasing, tigers drive leopards off to areas located closer to human settlements, like in Nepal's Bardia National Park and Sariska Tiger Reserve. Resource partitioning occurs where leopards share their range with tigers. Leopards tend to take smaller prey, usually less than , where tigers are present. In areas where leopard and tiger are sympatric, coexistence is reportedly not the general rule, with leopards being few where tigers are numerous. The mean leopard density decreased significantly (from 9.76 to 2.07 animals per ) while the mean density of tigers increased (from 3.31 to 5.81 animals/100 km2) from 2004–2005 to 2008 in Rajaji National Park following the relocation of pastoralists out of the park. There, the two species have high dietary overlap, and an increase in the tiger population resulted in a sharp decrease in the leopard population and a shift in the leopard diet to small prey (from 9% to 36%) and domestic prey (from 6.8% to 31.8%). In Chitwan National Park, leopards killed prey ranging from less than in weight with most kills in the range. Tigers killed more prey in the range. There were also differences in the microhabitat preferences of the individual tiger and leopard followed over five months; the tiger used roads and forested areas more frequently, while the leopard used recently burned areas and open areas more frequently. When a tiger killed baits at sites formerly frequented by leopards, the leopards did not hunt there for some time. In the tropical forests of India's
Nagarhole National Park Nagarahole National Park is a national park located in Kodagu district and Mysore district in Karnataka, India. This park was declared the 37th Tiger Reserve of India in 1999. It is part of the Nilgiri Biosphere Reserve. The Western Ghats ...
, tigers selected prey weighing more than , whereas leopards selected prey in the range. In tropical forests, they do not always avoid the larger cats by hunting at different times. With relatively abundant prey and differences in the size of prey selected, tigers and leopards seem to successfully coexist without competitive exclusion or interspecies dominance hierarchies that may be more common to the leopard's co-existence with the lion in savanna habitats. In areas with high tiger populations, such as in the central parts of India's
Kanha National Park Kanha Tiger Reserve, also known as Kanha–Kisli National Park, is one of the tiger reserves of India and the largest national park of the state of Madhya Pradesh. The present-day Kanha area is divided into two protected areas, Hallon and Banjar ...
, leopards are not permanent residents, but transients. They were common near villages at the periphery of the park and outside the park. In a reserved forest of southern India, species preyed upon by leopard, dhole and striped hyena overlapped considerably. The leopard and
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...
both hunt
Himalayan tahr The Himalayan tahr (''Hemitragus jemlahicus'') is a large even-toed ungulate native to the Himalayas in southern Tibet, northern India, western Bhutan and Nepal. It is listed as Near Threatened on the IUCN Red List, as the population is de ...
and
musk deer Musk deer can refer to any one, or all seven, of the species that make up ''Moschus'', the only extant genus of the family Moschidae. Despite being commonly called deer, they are not true deer belonging to the family Cervidae, but rather their f ...
, but the leopard usually prefers forested habitats located at lower altitudes. Leopard may conflict with
sloth bears The sloth bear (''Melursus ursinus'') is a myrmecophagous bear species native to the Indian subcontinent. It feeds on fruits, ants and termites. It is listed as vulnerable on the IUCN Red List, mainly because of habitat loss and degradation. ...
and can follow them up trees. Bear cubs are probably far more vulnerable and healthy adult bears may be avoided by leopards. One leopard killed a three-quarters grown female sloth bear in an apparently lengthy fight that culminated in the trees. Apparently, a sloth bear killed a leopard in a confrontation in Yala National Park,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
but was itself badly injured in the fight and was subsequently put down by park rangers.


Threats

Hunting of Indian leopards for the illegal
wildlife trade Wildlife trade refers to the of products that are derived from non-domesticated animals or plants usually extracted from their natural environment or raised under controlled conditions. It can involve the trade of living or dead individuals, t ...
is the biggest threat to their survival. They are also threatened by loss of habitat and fragmentation of formerly connected populations, and various levels of human–leopard conflict in human–dominated landscapes. Several newspapers reported of leopards falling into open wells and being rescued with the help of Forest Department officials.


Poaching

A significant immediate threat to wild leopard populations is the illegal trade in poached skins and body parts between India, Nepal and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. The governments of these countries have failed to implement adequate enforcement response, and wildlife crime remained a low priority in terms of political commitment and investment for years. There are well-organised gangs of professional poachers, who move from place to place and set up camp in vulnerable areas. Skins are rough-cured in the field and handed over to dealers, who send them for further treatment to Indian tanning centres. Buyers choose the skins from dealers or tanneries and smuggle them through a complex interlinking network to markets outside India, mainly in China.Banks, D., Lawson, S., Wright, B. (eds.) (2006)
''Skinning the Cat: Crime and Politics of the Big Cat Skin Trade''
Environmental Investigation Agency, Wildlife Protection Society of India
Seized skins in
Kathmandu , pushpin_map = Nepal Bagmati Province#Nepal#Asia , coordinates = , subdivision_type = Country , subdivision_name = , subdivision_type1 = Province , subdivision_name1 = Bagmati Prov ...
confirm the city's role as a key staging point for illegal skins smuggled from India bound for Tibet and China.Banks, D. (2004)
''The Tiger Skin Trail''
Environmental Investigation Agency.
It is likely that seizures represent a tiny fraction of the total illegal trade, with the majority of smuggled skins reaching their intended end market. Seizures revealed: * in India: more than 200 leopards killed by humans every year, leopards in India is 7 times more likely to be killed than Indian tigers. WPSI reported that during 1994-2010 at least 3,189 leopards were killed, then again in 2002-2010 period at least 200 or four leopards per week were reportedly killed by poachers for illegal trade. For every
tiger The tiger (''Panthera tigris'') is the largest living Felidae, cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily pr ...
skin, there are at least seven leopard skins in the haul.Wildlife Protection Society of India (2010)
''Leopards Battling For Survival In India''
Wildlife Protection Society of India, 18 May 2010.
* in Nepal: more than 40 leopards were reported killed by humans every year, e.g. 243 poached leopards between May 2002 and May 2008; * in China and
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
: nearly 130 leopards were killed every year, e.g. more than 774 poached leopards between July 1999 and September 2005.


Human–leopard conflict


Causes of conflict

Expansion of agriculturally used land, encroachment by humans and their livestock into protected areas are main factors contributing to habitat loss and decrease of wild prey. As a result, leopards approach human settlements, where they are tempted to prey on dogs, pigs and goats – domestic livestock, which constitutes an important part of their diet, if they live on the periphery of human habitations. Human–leopard conflict situations ensue, and have increased in recent years. In retaliation for attacks on livestock, leopards are shot, poisoned and trapped in snares. The leopards are considered to be unwanted trespassers by villagers. Conservationists criticize these actions, claiming that people are encroaching on the leopard's native habitat. India's Forest Department is entitled to set up traps only in cases of a leopard having attacked humans. If only the presence of a crowd of people prevents the leopard from escaping, then the crowd has to be dispersed and the animal allowed to escape. As urban areas expanded, the natural habitats of leopards shrunk resulting in leopards venturing into urbanized areas due to easy access of domestic food sources. Karnataka has a high number of such conflicts. In recent years, leopards were sighted in
Bangalore Bangalore (), officially Bengaluru (), is the capital and largest city of the Indian state of Karnataka. It has a population of more than and a metropolitan population of around , making it the third most populous city and fifth most ...
, and the forest department captured six leopards in the city's outskirts, relocated four of them to various other locations.


Man-eater leopards

Every year more leopards are killed by humans than the humans killed by leopards. On average nearly 400 leopards are reported killed yearly in India, Nepal and China combined based on the leopard skins caught from the poachers, though the actual number of leopards killed by humans is likely to be several times higher. In and around the
Shivalik hills The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Ind ...
of
Himachal Pradesh Himachal Pradesh (; ; "Snow-laden Mountain Province") is a state in the northern part of India. Situated in the Western Himalayas, it is one of the thirteen mountain states and is characterized by an extreme landscape featuring several pea ...
alone, 68 leopards were killed by people between 2001 and 2013, of which only 10 were man-eaters. The frequency of Leopard attacks on humans varies by geographical region and historical period. Since India and
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
have the majority of Indian leopards population, consequently attacks are regularly reported only from India and Nepal. Among the five "
big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. Despite enormous differences in size, various cat species are quite similar ...
s", leopards are less likely to become
man-eater A man-eater is an animal that preys on humans as a pattern of hunting behavior. This does not include the scavenging of corpses, a single attack born of opportunity or desperate hunger, or the incidental eating of a human that the animal has kil ...
s—only
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus ''Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
s and
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...
s have a less fearsome reputation. While leopards generally avoid humans, they tolerate proximity to humans better than
lion The lion (''Panthera leo'') is a large cat of the genus '' Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adu ...
s and
tiger The tiger (''Panthera tigris'') is the largest living Felidae, cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily pr ...
s and often come into conflict with humans when raiding livestock. Attacks in India are still reported, since leopards population in India outnumber population of all other large carnivores combined, consequently the number of humans killed by leopards is also more than those killed by all other large carnivores combined. In Nepal, where most attacks occur in the midland regions, i.e. in the
Terai , image =Terai nepal.jpg , image_size = , image_alt = , caption =Aerial view of Terai plains near Biratnagar, Nepal , map = , map_size = , map_alt = , map_caption = , biogeographic_realm = Indomalayan realm , global200 = Terai-Duar savanna a ...
, midhills, and
lesser Himalaya The Lower Himalayan Range ( ne, पर्वत शृङ्खला parbat shrinkhalā) – also called the Middle Himalayas or Lesser Himalayas or Himachal – is a major east–west mountain range with elevations 3,700 to 4,500 m (12,000 to ...
, the rate of leopard predation on humans results in approximately 1.9 human deaths annually per million inhabitants. . Historically, with rapid urbanization in late 19th and early 20th centuries, leopard attacks may have peaked in India during those times. Notable man-eaters of that era include Leopard of
Central Provinces The Central Provinces was a province of British India. It comprised British conquests from the Mughals and Marathas in central India, and covered parts of present-day Madhya Pradesh, Chhattisgarh and Maharashtra states. Its capital was Nagpur ...
, Rudraprayag, Gummalapur, Yellagiri Hills, Golis Range and Panar.


Ways to minimise conflict

Key to avoiding conflict or leopard's predation of humans is to shift the focus on human's behavioral change to minimise the chances of a leopard encounter or attack, which can be achieved by "clearing bushes and overgrowth around homes to minimise hiding spaces for leopards, leaving a light on at night to deter them, and ensuring people, especially children, did not go out alone at night." Leopards are shy and avoid humans and are more active at night, during encounter with leopards "give way to the leopard and move away calmly" and alert the forest department immediately.Spotted in India: Humans and leopards living in harmony
ourbetterworld.org, 9 Apr 2021.


Conservation

''Panthera pardus'' is listed in
CITES Appendix I CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
. Despite India and Nepal being contracting parties to
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
, national legislation of both countries does not incorporate and address the spirit and concerns of CITES. Trained human resources, basic facilities and effective networks for control of poaching and trade in wildlife are lacking. The Indian leopard is considered Vulnerable in India, Bhutan, and Nepal but Critically Endangered in Pakistan. Frederick Walter Champion was one of the first in India who after
World War I World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was List of wars and anthropogenic disasters by death toll, one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, ...
advocated for the conservation of leopards, condemned sport hunting and recognised their key role in the ecosystem. Billy Arjan Singh championed their cause since the early 1970s. There a few leopard rescue centres in India, such as the Manikdoh Leopard Rescue Centre in
Junnar Junnar (Marathi pronunciation: ͡ʒunːəɾ is a city in the Pune district of the Indian state of Maharashtra. The city has history dating back to the first millennium. The nearby fort of Shivneri was the birthplace of Maratha king Chatrap ...
, but more rescue and rehabilitation centres are being planned. Some wildlife experts think that such centres are not an ideal solution, but that conflict resolution by way of changing human behaviour, land use or grazing patterns and implementing responsible forest management to lessen human-animal conflict would be far more effective to conserve leopards.


In culture and literature

*An Indian
black panther A black panther is the melanistic colour variant of the leopard (''Panthera pardus'') and the jaguar (''Panthera onca''). Black panthers of both species have excess black pigments, but their typical rosettes are also present. They have been ...
named '
Bagheera Bagheera ( hi, बघीरा / ''Baghīrā'') is a fictional character in Rudyard Kipling's Mowgli stories in ''The Jungle Book'' (coll. 1894) and ''The Second Jungle Book'' (coll. 1895). He is a black panther (melanistic Indian leopard) who ...
' is featured in
Rudyard Kipling Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)'' The Times'', (London) 18 January 1936, p. 12. was an English novelist, short-story writer, poet, and journalist. He was born in British India, which inspired much of his work. ...
's 1894 novel ''
The Jungle Book ''The Jungle Book'' (1894) is a collection of stories by the English author Rudyard Kipling. Most of the characters are animals such as Shere Khan the tiger and Baloo the bear, though a principal character is the boy or "man-cub" Mowgli, w ...
'', as well as in
Disney The Walt Disney Company, commonly known as Disney (), is an American multinational mass media and entertainment conglomerate headquartered at the Walt Disney Studios complex in Burbank, California. Disney was originally founded on October ...
's
1967 Events January * January 1 – Canada begins a year-long celebration of the 100th anniversary of Confederation, featuring the Expo 67 World's Fair. * January 5 ** Spain and Romania sign an agreement in Paris, establishing full consular and ...
and
2016 File:2016 Events Collage.png, From top left, clockwise: Bombed-out buildings in Ankara following the 2016 Turkish coup d'état attempt; the Impeachment of Dilma Rousseff, impeachment trial of Brazilian President Dilma Rousseff; Damaged houses duri ...
film adaptations. * The book '' Man-Eaters of Kumaon'' is based on man-eating leopards and tigers in
Kumaon Kumaon or Kumaun may refer to: * Kumaon division, a region in Uttarakhand, India * Kumaon Kingdom, a former country in Uttarakhand, India * Kumaon, Iran, a village in Isfahan Province, Iran * , a ship of the Royal Indian Navy during WWII See also ...
.''Man-Eaters of Kumaon'' (1944), Jim Corbett,
Oxford University Press Oxford University Press (OUP) is the university press of the University of Oxford. It is the largest university press in the world, and its printing history dates back to the 1480s. Having been officially granted the legal right to print book ...
,
Bombay Mumbai (, ; also known as Bombay — List of renamed Indian cities and states#Maharashtra, the official name until 1995) is the capital city of the Indian States and union territories of India, state of Maharashtra and the ''de facto'' fin ...
.
* ''
Ajoba Ajoba is a 2014 Indian Marathi-language film directed by Sujay Dahake and written by Gauri Bapat. It is allegedly based on true events and relates to how a leopard trekked from Malshej Ghat to Mumbai Mumbai (, ; also known as Bombay � ...
'' is a 2014 Marathi film directed by
Sujay Dahake Sujay Sunil Dahake (Marathi सुजय सुनील डहाके) is mainly known for his work as director and as editor for the film ''Shala Shala (Šala) was a Mesopotamian goddess of weather and grain and the wife of the weather god ...
and written by Gauri Bapat. It is supposedly based on true events. * Local people in the Western Ghats call a darkish grey panther 'Pogeyan', which purportedly lives in the high-altitude grasslands of the Anamudi forest.


See also

* Leopard subspecies * Chinese leopard *
Zanzibar leopard The Zanzibar leopard is an African leopard (''Panthera pardus pardus'') population on Unguja Island in the Zanzibar archipelago, Tanzania, that is considered extirpated due to persecution by local hunters and loss of habitat. It was the island's ...


References


Further reading

* *


External links

* Species portrai
''Panthera pardus'' in Asia
and short portrai
''P. pardus fusca''
IUCN/SSC Cat Specialist Group
wild-cat.org — Information about research and conservation of leopards in Asia

''Project Waghoba'' — Leopard Research and Conservation Project in Western Maharashtra, India

Wildlife SOS: ''After 5 years, 3 leopards get a better life''


{{Taxonbar, from=Q729553
Leopards The leopard (''Panthera pardus'') is one of the five extant species in the genus ''Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
Felids of India Fauna of South Asia Mammals of Pakistan Mammals of Nepal Mammals of India Mammals of Bhutan Mammals of Bangladesh Indian leopard