''Isatis'' is a genus of
flowering plants in the family
Brassicaceae, native to the
Mediterranean region east to
central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes
woad (''Isatis tinctoria''). Due to their extremely variable morphology, the Asian species in particular are difficult to determine; the only reliable diagnostic feature is the ripe fruit. They are (usually)
biennial or perennial
herbaceous plants, often bluish and hairless or downy hairy with the upright stem branched.
Description
They are annual, biennial or perennial, branched herbs, usually
glabrous and
glaucous
''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), g ...
except
silique. Basal leaves generally elliptic-oblong,
sessile; sessile caulinary, rounded to oval-oblong.
The
hermaphrodite
In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes.
Many Taxonomy (biology), taxonomic groups of animals (mostly invertebrate ...
flowers are fourfold double
perianth. The four
sepals are ascending to upright. The four yellow to
off-white
Shades of white are colors that differ only slightly from pure white. Variations of white include what are commonly termed off-white colors, which may be considered part of a neutral color scheme or yellow that looks like brown.
In color theo ...
or
lilac
''Syringa'' is a genus of 12 currently recognized species of flowering plant, flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and wid ...
-white petals are at least as long as the sepals. They have six
stamen
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filame ...
s with very small, egg-shaped or elongated-round anthers. There are
nectar
Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
glands.
Racemose is branched or paniculated, ebracted, inflorescence, often reaching lax and elongated in the fruit.
The fruit is a generally linear silique, oblong-cuneate to suborbicular, indehiscent, flattened laterally, unilocular, little to conspicuously winged, glabrous or with tiny hairs.
[Flora of North America Editorial Committee, e. 2010. Magnoliophyta: Salicaceae to Brassicaceae. Fl. N. Amer. 7: i – xxii, 1–797.]
Species
Currently accepted species include:
*''
Isatis afghanica''
Hadac & Chrtek
*''
Isatis amani''
P.H.Davis
*''
Isatis apennina''
Ten. ex Grande
*''
Isatis apscheronica''
N.Busch
*''
Isatis aptera''
(Boiss. & Heldr.) Al-Shehbaz, Moazzeni & Mumm.
*''
Isatis arenaria''
Azn.
*''
Isatis armena''
L.
*''
Isatis arnoldiana''
N.Busch
*''
Isatis aucheri''
Boiss.
*''
Isatis biscutellifolia''
Boiss. & Buhse
*''
Isatis bitlisica''
P.H.Davis
*''
Isatis boissieriana
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Rchb.f.
*''
Isatis brachycarpa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''Is ...
''
C.A.Mey.
*''
Isatis brevipes''
(Bunge) Jafri
*''
Isatis bullata''
Aitch. & Hemsl.
*''
Isatis bungeana''
Seidlitz
*''
Isatis buschiana
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Schischk.
*''
Isatis callifera
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''Is ...
''
Boiss. & Balansa
*''
Isatis campylocarpa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Boiss.
*''
Isatis canaliculata
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
(Vassilcz.) V.V.Botschantz.
*''
Isatis candolleana
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Boiss.
*''
Isatis cappadocica''
Desv.
*''
Isatis cardiocarpa''
(Trautv.) Al-Shehbaz, Moazzeni & Mumm.
*''
Isatis caucasica
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
N.Busch
*''
Isatis cochlearis
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Boiss.
*''
Isatis constricta
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''Is ...
''
P.H.Davis
*''
Isatis costata''
C.A.Mey.
*''
Isatis davisiana
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
H.Misirdali ex P.H.Davis, R.R.Mill & Kit Tan
*''
Isatis demiriziana''
H.Misirdali ex P.H.Davis, R.R.Mill & Kit Tan
*''
Isatis densiflora''
(Bunge ex Boiss.) D.A.German
*''
Isatis deserti''
(N.Busch) V.V.Botschantz.
*''
Isatis djurjaedae
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Coss. & Durieu
*''
Isatis elegans
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
(Boiss.) Hadac & Chrtek
*''
Isatis emarginata''
Kar. & Kir.
*''
Isatis erzurumica''
P.H.Davis
*''
Isatis floribunda
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Boiss. ex Bornm.
*''
Isatis frigida''
Boiss. & Kotschy
*''
Isatis frutescens
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Kar. & Kir.
*''
Isatis gaubae
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Bornm.
*''
Isatis glastifolia''
(Fisch. & C.A.Mey.) Al-Shehbaz, Moazzeni & Mumm.
*''
Isatis glauca
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Aucher ex Boiss.
*''
Isatis grammotis
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Kit Tan
*''
Isatis gymnocarpa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''Is ...
''
(Fisch. ex DC.) Al-Shehbaz, Moazzeni & Mumm.
*''
Isatis harsukhii''
O.E.Schulz
*''
Isatis hirtocalyx''
Franch.
*''
Isatis huber-morathii''
P.H.Davis
*''
Isatis iberica''
Steven
*''
Isatis jacutensis
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
(N.Busch) N.Busch
*''
Isatis karjaginii''
Schischk.
*''
Isatis kotschyana''
Boiss. & Hohen.
*''
Isatis kozlovskyi
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Grossh.
*''
Isatis laevigata
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Trautv.
*''
Isatis latisiliqua''
Steven
*''
Isatis leuconeura
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Boiss. & Buhse
*''
Isatis littoralis''
Steven
*''
Isatis lockmanniana''
Kotschy ex Boiss.
*''
Isatis lusitanica
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
L.
*''
Isatis mardinensis''
P.H.Davis & H.Misirdali
*''
Isatis maxima
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Pavlov
*''
Isatis microcarpa''
J.Gay ex Boiss.
*''
Isatis minima
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Bunge
*''
Isatis multicaulis
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
(Kar. & Kir.) Jafri
*''
Isatis oblongata
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
DC.
*''
Isatis odontogera''
(Bordz.) D.A.German
*''
Isatis ornithorhynchus''
N.Busch
*''
Isatis pachycarpa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Rech.f., Aellen & Esfand.
*''
Isatis pinnatiloba''
P.H.Davis
*''
Isatis platyloba
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Link ex Steud.
*''
Isatis praecox''
Kit. ex Tratt.
*''
Isatis raimondoi
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Di Grist., Scafidi & Domina
*''
Isatis raphanifolia
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Boiss.
*''
Isatis rugulosa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Bunge ex Boiss.
*''
Isatis sabulosa
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Steven ex Ledeb.
*''
Isatis sevangensis''
N.Busch
*''
Isatis sivasica
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
P.H.Davis
*''
Isatis spatella''
P.H.Davis
*''
Isatis spectabilis
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
P.H.Davis
*''
Isatis steveniana
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Trautv.
*''
Isatis stocksii
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Boiss.
*''
Isatis subdidyma''
(N.Busch) V.E.Avet.
*''
Isatis takhtajanii''
Avet.
*''
Isatis tinctoria''
L.
*''
Isatis tomentella''
Boiss. & Balansa
*''
Isatis trachycarpa''
Trautv.
*''
Isatis turcomanica''
Korsh.
*''
Isatis undulata
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad (''I ...
''
Aucher ex Boiss.
*''
Isatis violascens
''Isatis'' is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. Its genus name, Isatis, derives from the ancient Greek word for the plant, ἰσάτις. The genus includes woad ...
''
Bunge
*''
Isatis zarrei''
Al-Shehbaz, Moazzeni & Mumm.
References
Brassicaceae
Brassicaceae genera
{{Brassicales-stub