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''Isaberrysaura'' is a genus of
The holotype specimen of ''Isaberrysaura'' was discovered in the marine to deltaic sediments of the Los Molles Formation, which is of Bajocian age, by Isabel Valdivia Berry, who is honoured in the generic name. The holotype, known as MOZ-Pv 6459 consists of a partial skeleton with a nearly complete skull (the post-cranial material of which is still unprepared), six cervical vertebrae, fifteen
The holotype specimen of ''Isaberrysaura'' has been preserved with fossilized stomach contents. Within the rib cage, a mass of fossilized seeds was discovered; the first preserved meal uncovered in a basal ornithischian. Two types of seeds were recovered close to the posterior ribs of ''Isaberrysaura'', distinguished according to their size. The largest seeds preserved three layers: an outer fleshy sarcotesta, the sclerotesta, and the inner layer (possibly corresponding to the nucellus). These seeds are cycadales of the family Zamiineae on the basis of a well-defined coronula in the micropylar region, whereas the smaller, platyspermic seeds are still indeterminate. The intact nature of the sarcotesta in the largest seeds suggests that ''Isaberrysaura'' swallowed the seeds whole without chewing and that they were in the first stages of digestion based on their position in the gut. Based on its teeth, it is possible that the dinosaur had an omnivorous diet, but as no trace of animal remains have been found alongside the seeds, this remains speculative at best. It also remains possible that there were enzyme-producing bacteria present in the dinosaur's gut to aid in digesting tougher plant material as seeds.
stegosauria
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, ...
n ornithischian dinosaur from the Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
Los Molles Formation of Patagonia, Argentina. The genus contains a single species, ''I. mollensis'', described by Salgado ''et al.'' in 2017 from a single specimen. Although initially classified as a basal neornithischian, subsequent analysis has allied it with the Stegosauria
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, ...
; the morphology of its skull resembles those of other members of the group.
Description
The teeth of ''Isaberrysaura'' areheterodont
In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology.
In vertebrates, heterodont pertains to animals where teeth are differentiated into different forms. For example, ...
, with recurved premaxillary teeth and lanceolate maxillary and dentary teeth. In extant iguanid lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s, similar dentition is correlated with omnivorous diets, indicating that ''Isaberrysaura'' might too have been omnivorous. ''Isaberrysaura'' is also estimated to have measured around long, making it of moderate size.
The skull in particular is very unusual; it is estimated to be 52 cm long and 20 cm wide across the orbits, and it is almost as high as it is wide. The snout slopes anteroventrally from the posterodorsal corner of the infratemporal fenestra to what is seemingly the maxillary-premaxillary contact and the infratemporal fenestra is dorsoventrally deep. In contrast, the orbit is subcircular, not quite as dorsoventrally tall as anteroposteriorly long, smaller than the infratemporal fenestra, and only visible in lateral view. The anterolateral sector of the left supratemporal fenestra is relatively well preserved and it is visible only in dorsal view. The skull's antorbital fossa is roughly triangular, with its base longer than the other two sides. The jugal is triradiate and the anterior process of the jugal forms the posteroventral corner of the antorbital fossa and surpasses anteriorly the base of the lacrimal, a feature seen in basal thyreophorans and stegosaurs
Stegosauria is a group of Herbivore, herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous Period (geology), periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what i ...
. The dorsal process of the jugal is proportionally long. The quadratojugal is very broad and the premaxilla is incompletely preserved while the post-cranial material is as-yet-undescribed at present. The dentition is heterodont, with six premaxillary teeth and thirty maxillary teeth. The genus' overall appearance is highly similar to that of stegosaurians, though lacking the characteristic plates and likely possessing a bipedal stance.
Discovery
The holotype specimen of ''Isaberrysaura'' was discovered in the marine to deltaic sediments of the Los Molles Formation, which is of Bajocian age, by Isabel Valdivia Berry, who is honoured in the generic name. The holotype, known as MOZ-Pv 6459 consists of a partial skeleton with a nearly complete skull (the post-cranial material of which is still unprepared), six cervical vertebrae, fifteen dorsal vertebra
In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebrae and they are intermediate in size between the cervical ...
e, a sacrum
The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30.
The sacrum situates at the upper, back part ...
with a partial ilium
Ilium or Ileum may refer to:
Places and jurisdictions
* Ilion (Asia Minor), former name of Troy
* Ilium (Epirus), an ancient city in Epirus, Greece
* Ilium, ancient name of Cestria (Epirus), an ancient city in Epirus, Greece
* Ilium Building, a ...
and an apparently complete pubis, nine caudal vertebrae, part of a scapula, ribs, with the addition of unidentifiable fragments.
Classification
''Isaberrysaura's'' classification is problematic at best; the holotype specimen shares traits with both thyreophorans and neornithischians. For this reason, it has been placed in the Genasauria. Three datasets within the same study by Salgado ''et al'' indicated that while it possessed the characteristics of the neornithischians and thyreophorans, it did not fit within either of those groups, representing a previously unknown morphotype within the Ornithischia as a whole. Subsequently, the phylogenetic analysis conducted by Han ''et al.'' (2018) recovered ''Isaberrysaura'' as one of the basalmost known stegosaurs.Paleobiology
The holotype specimen of ''Isaberrysaura'' has been preserved with fossilized stomach contents. Within the rib cage, a mass of fossilized seeds was discovered; the first preserved meal uncovered in a basal ornithischian. Two types of seeds were recovered close to the posterior ribs of ''Isaberrysaura'', distinguished according to their size. The largest seeds preserved three layers: an outer fleshy sarcotesta, the sclerotesta, and the inner layer (possibly corresponding to the nucellus). These seeds are cycadales of the family Zamiineae on the basis of a well-defined coronula in the micropylar region, whereas the smaller, platyspermic seeds are still indeterminate. The intact nature of the sarcotesta in the largest seeds suggests that ''Isaberrysaura'' swallowed the seeds whole without chewing and that they were in the first stages of digestion based on their position in the gut. Based on its teeth, it is possible that the dinosaur had an omnivorous diet, but as no trace of animal remains have been found alongside the seeds, this remains speculative at best. It also remains possible that there were enzyme-producing bacteria present in the dinosaur's gut to aid in digesting tougher plant material as seeds.
See also
*2017 in archosaur paleontology
The year 2017 in archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology ...
References
{{Taxonbar, from=Q28791257 Thyreophorans Ornithischian genera Bajocian life Middle Jurassic dinosaurs of South America Jurassic Argentina Fossils of Argentina Neuquén Basin Fossil taxa described in 2017†
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...