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The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The ge ...
that is largely adapted to polar and mountainous habitats.


Evolution

The mountain hare arose during the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
; there is evidence that its range expanded during glaciations into southern Europe, with populations of
Iberian hare The Granada hare (''Lepus granatensis''), also known as the Iberian hare, is a hare species that can be found on the Iberian Peninsula and on the island of Majorca. Subspecies Three subspecies of the Granada hare are known, which vary in colour ...
(''Lepus granatensis''),
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
(''L. europaeus)'' and
broom hare The broom hare (''Lepus castroviejoi'') is a species of hare endemic to northern Spain. Distribution and habitat It is restricted to the Cantabrian Mountains in northern Spain between the Serra dos Ancares and the Sierra de Peña Labra. This ...
(''L. castroviejoi'') in northern Iberia harboring
mitochondrial A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is use ...
haplotype A haplotype ( haploid genotype) is a group of alleles in an organism that are inherited together from a single parent. Many organisms contain genetic material ( DNA) which is inherited from two parents. Normally these organisms have their DNA or ...
s from the mountain hare. During the Late Pleistocene to
Early Holocene In the geologic time scale, the Greenlandian is the earliest age or lowest stage of the Holocene Epoch or Series, part of the Quaternary. Beginning in 11,650 BP (9701 BCE or 300 HE) and ending 8,276 BP (6237 BCE or 3764 HE), it is the earlie ...
, populations of mountain hare in Russia grew at least 10% larger than any living population today. This population has been classified as a distinct species ''Lepus tanaiticus'', but is now generally considered a prehistoric
morphotype In biology, polymorphism is the occurrence of two or more clearly different morphs or forms, also referred to as alternative ''phenotypes'', in the population of a species. To be classified as such, morphs must occupy the same habitat at the s ...
of the living mountain hare.


Distribution

This species is distributed from
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
; in addition, isolated mountain populations occur in the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
,
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
, the
Baltics The Baltic states, et, Balti riigid or the Baltic countries is a geopolitical term, which currently is used to group three countries: Estonia, Latvia, and Lithuania. All three countries are members of NATO, the European Union, the Eurozone, ...
, northeastern
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
, and
Hokkaidō is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel. The la ...
. In
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
, the Irish hare (''L. t. hibernicus'') lives on lowland pastures, coastal grasslands, and salt marshes, not just in the mountains. The mountain hare has also been introduced to
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...
,
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
,
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
, the
Isle of Man ) , anthem = "O Land of Our Birth" , image = Isle of Man by Sentinel-2.jpg , image_map = Europe-Isle_of_Man.svg , mapsize = , map_alt = Location of the Isle of Man in Europe , map_caption = Location of the Isle of Man (green) in Europe ...
, the
Peak District The Peak District is an upland area in England at the southern end of the Pennines. Mostly in Derbyshire Derbyshire ( ) is a ceremonial county in the East Midlands, England. It includes much of the Peak District National Park, the southe ...
,
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
,
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...
,
Crozet Islands The Crozet Islands (french: Îles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarcti ...
, and the
Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...
. In the Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on geographic region and season.


Description

The mountain hare is a large species, though it is slightly smaller than the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
. It grows to a length of , with a tail of , and a mass of , females being slightly heavier than males. They can live for up to 12 years. In summer, for all populations of mountain hares, the coat is various shades of brown. In preparation for winter most populations moult into a white (or largely white)
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
. The tail remains completely white all year round, distinguishing the mountain hare from the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
(''Lepus europaeus''), which has a black upper side to the tail. The subspecies ''Lepus timidus hibernicus'' (the Irish mountain hare) is smaller in size and stays brown all year. The Irish hare may also have a "golden" variation, particularly those found on
Rathlin Island Rathlin Island ( ga, Reachlainn, ; Local Irish dialect: ''Reachraidh'', ; Scots: ''Racherie'') is an island and civil parish off the coast of County Antrim (of which it is part) in Northern Ireland. It is Northern Ireland's northernmost point. ...
. In the Faroe Islands, mountain hares turn grey in the winter instead of white. The winter-grey colour may be caused by downregulation of the agouti hair cycle isoform in the autumn moult.


Behaviour

The diet of the mountain hare varies from region to region. It seems to be somewhat dependent on the particular habitat in which the population under study lives. For example, in northern Scandinavia, where snow may blanket the ground for many months, the hares may feed on twigs and bark. In areas where snowfall is rare, such as Ireland,
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
may form the bulk of the diet. Given a choice, mountain hares in Scotland and Ireland seem to prefer grazing (feeding on grasses). For mountain hares on a coastal
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
environment in Ireland, grasses constituted over 90% of their diet. This was higher than the percentage of grass in the diet of the
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has ...
(''Oryctolagus cuniculus'') that inhabited the same environment. The mountain hare is regionally the favourite prey of the
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
and may additionally be preyed on by
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Palearctic, Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, ...
s and
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
es.
Stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
s may prey on young hares. In northern parts of Finland, Norway, and Sweden, the mountain hare and the European hare compete for habitat. The European hare, being larger, is usually able to drive away the mountain hare, but is less adapted for living in snowy regions; its feet are smaller and its winter fur is a mixture of white and brown. While this winter fur is actually a very good camouflage in the coastal regions of Finland where the snow covers the shrubs only a short time, the mountain hare is better adapted for the snowier conditions of the inland areas. The two may occasionally cross. The
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and ...
(''Lepus arcticus'') was once considered a subspecies of the mountain hare, but it is now regarded as a separate species. Similarly, some scientists believe that the Irish hare should be regarded as a separate species. Fifteen subspecies are currently recognised.


Human impact

In the European Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on biographic region and season. The development of alpine winter tourism has increased rapidly since the last few decades of the 20th century, resulting in expansion of ski resorts, growing visitor numbers, and a huge increase in all forms of snow sport activities. A 2013 study looking at stress events and the response of mountain hares to disturbance concluded that those hares living in areas of high winter recreational activities showed changes in physiology and behaviour that demanded additional energy input at a time when access to food resources is restricted by snow. It recommended ensuring that forests inhabited by mountain hares were kept free of tourist development, and that new skiing areas should be avoided in mountain hare habitat, and that existing sites should not be expanded. In August 2016, the Scottish animal welfare charity
OneKind OneKind is a Advocacy group, campaigning animal welfare charity based in Edinburgh and operating in Scotland, UK and as part of the Eurogroup for Animals. OneKind exists to end cruelty to all animals and improve their lives. Background The org ...
launched a campaign on behalf of the mountain hare, as a way of raising awareness of mountain hare culls taking place across the country and in garnering public support for the issue. Mountain hares are routinely shot in the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland Sco ...
both as part of paid hunting "tours" and by gamekeepers managing red grouse populations (who believe that mountain hares can be vectors of diseases that affect the birds). Much of this activity is secretive, but investigations have revealed that tens of thousands of hares are being culled every year. The campaign, which urges people to proclaim, "We Care For The Mountain Hare", will culminate with the charity urging the Scottish government to legislate against commercial hunting and culling of the iconic Scottish species. The campaign has revealed widespread public support for a ban on hare hunting in Scotland. On May 17, 2020,
MSPs Member of the Scottish Parliament (MSP; gd, Ball Pàrlamaid na h-Alba, BPA; sco, Memmer o the Scots Pairliament, MSP) is the title given to any one of the 129 individuals elected to serve in the Scottish Parliament. Electoral system The ad ...
voted to ban the unlicensed culling of mountain hares and grant them protected species status within Scotland after a petition started by Green MSP
Alison Johnstone Alison Johnstone (born 11 October 1965) is a Scottish politician who has served as the Presiding Officer of the Scottish Parliament since 2021. Elected as a member of the Scottish Greens, she relinquished her party affiliation on becoming Presi ...
gathered over 22,000 signatures. In 2021, the
People's Trust for Endangered Species Peoples Trust for Endangered Species (PTES) is a charitable organisation registered in England and Wales. It exists to promote the conservation of rare or declining species and habitats in the UK and worldwide through environmental monitoring, m ...
funded a survey of mountain hare populations in the UK's Peak District after concerns about the viability of the isolated population, believed to be as low as 2,500. The trust believes climate change is a threat to long-term survival of the Peak District population, which was introduced to the area in the 1870s.


References


External links


Irish Hare Initiative
{{Taxonbar, from=Q180035 Lepus Mammals of Asia Mammals of Europe Mammals of China Mammals of Japan Mammals of Central Asia Mammals of Russia Arctic land animals Mammals described in 1758 Taxa named by Carl Linnaeus National symbols of the Republic of Ireland Mammals of Mongolia