Iris Sambucina
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''Iris sambucina'', the elder scented iris, is a plant species in the genus '' Iris'', it is also in the subgenus '' Iris''. It is a rhizomatous
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, from southern and central Europe (
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
, GB and
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
, the
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
), and
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
. It has green, curved or sword-like leaves, tall round stem (with branches), multiple flowers in shades from brown violet, or brown-purple, to purple-violet, blue violet, mauve, and to purple. The large flowers are fragrant, with the scent of elderflowers, hence the name. It was first considered a separate species (when found), then it was classified as a synonym of '' Iris germanica'', before being classified (in some sources) as a separate species again, but with a hybrid origin from '' Iris pallida'' and ''
Iris variegata ''Iris variegata'', commonly known as the Hungarian iris, is a plant species in the genus ''Iris'', also in the subgenus ''Iris''. It is a rhizomatous perennial from eastern Europe. It has dark green, ribbed leaves. The branched flowering stems c ...
''. It is sometimes cultivated as an ornamental plant in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
regions.


Description

It is similar in form to ''Iris germanica'', but it has darker colour flowers and fragranced flowers. It has thick, fleshy or fibrous rhizomes, that are well branched. It has
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
, glaucous,William Robinson sword-like, or curving, green leaves. They can grow (in spring,) up to between long, and between 1.3 and 2.5 cm wide. They also smell strongly of elder ('' Sambucus ebulus''). It has a round stem, or
peduncle Peduncle may refer to: *Peduncle (botany), a stalk supporting an inflorescence, which is the part of the shoot of seed plants where flowers are formed *Peduncle (anatomy), a stem, through which a mass of tissue is attached to a body **Peduncle (art ...
, that can grow up to between tall. The stem is normally taller than the leaves,
Lady Charlotte Murray Lady Charlotte Murray (2 August 1754 – 4 April 1808) was a Scottish botanist and author. She was the eldest child of John Murray, 3rd Duke of Atholl, and Charlotte Murray, Duchess of Atholl. Her paternal grandfather was the Jacobite genera ...
and they are usually branched. The stem has large, spathes (leaves of the flower bud), that are green at the base but (scarious) or membranous on the top half. The stems (and the many branches) hold 2,William Prince or more flowers, blooming in summer, between April, or May, to June. The large, around long flowers, are fragrant (with the scent of elderflowers), and come in shades of violet, from brown violet, or brown-purple, purple-violet, blue violet, mauve, to purple. They are normally darker than ''Iris germanica''.Daniel Chambers Macreight They can also vary in colour from region to region. Like other irises, it has 2 pairs of petals, 3 large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
(outer petals), known as the 'falls' and 3 inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
), known as the 'standards'. The deflexed falls are spathulate (spoon-like) or oblong shaped, and can be veined, brown-purple on yellow-white background, or with claret (dark red). In the middle of the falls, is a row of short hairs called the 'beard', which is yellow, golden yellow, or orange. The upright standards are elliptical or oblong shaped, although the tips are inclined to each other. They are also serrated, or jagged. The style arms are light purple and 3–4 cm long, the perianth tube is 2–2.5 cm long. After the iris has flowered, most irises produces a seed capsule, but due to the hybrid origin, the capsules are not usually developed.


Genetics

As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
, having two sets of
chromosomes A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
, this can be used to identify hybrids and classification of groupings. It is published, that it a chromosome count: 2n=24, or 2n=40. As ''Iris germanica'' has a count of 2n=44 or 48 and ''Iris variegata'' is counted as 2n=24. ''Iris variegata'' is more likely to be the parent species.


Taxonomy

It is commonly known as 'elder scented iris', or 'Iris elderflower'. It is known as 'fläderiris' in
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
, or 'holunder schwertlilie' in Germany, (meaning 'elderberry iris'). The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''sambucina'' refers to sambuci-, sambucinus, -a, or -um elder-like or resembling Sambucus. This refers to the smell of the flowers resembling the scent of elder-flowers. It was first published and described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his book
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
(Syst. Nat) Edition 10, Volume 2 on page 863, on 7 June 1759. It was later named ''Iris glauca'' by Richard Anthony Salisbury in the ''
Transactions of the Horticultural Society of London ''The Garden'' is the monthly magazine of the British Royal Horticultural Society (RHS), circulated to all the society's members as a benefit of membership; it is also sold to the public. History ''The Garden'' magazine has gone under this title ...
'' Vol.1 on page 302 in 1812, but this was rejected by all other authors, due to Salisbury's plagiarism history. It was also published in Curtis's Botanical Magazine Vol.6 on tab.187 (with a colour illustration) in 1794, The Gardeners' Chronicle in 1883, the 'Handbook of the Irideae' by J. P. Baker, on page 86 in 1892 and the Journal of RHS Vol.15 on page 3 in October 1899.
William Rickatson Dykes William Rickatson Dykes (4 November 1877 – 1 December 1925) was an English amateur botanist who became an expert in the field of iris breeding and wrote several influential books on the subject. He was also interested in tulips, amaryllis, and ...
notes that Linnaeus had given the names of ''Iris sambucina'' and ''Iris squalens'', to 2 similar irises, whose parents were ''Iris pallida'' and ''Iris variegata''. It was later classified as a hybrid, but it is unsure which species (of iris) are the parents. It probably originated in the Mediterranean as a diploid species. Many authors think it is a hybrid of ''Iris variegata'' L. and ''Iris pallida'' Lam. Others think it is a synonym of ''Iris germanica'' L., including
The Plant List The Plant List was a list of botanical names of species of plants created by the Royal Botanic Gardens, Kew and the Missouri Botanical Garden and launched in 2010. It was intended to be a comprehensive record of all known names of plant species ...
. It is listed in the Encyclopedia of Life and
Catalogue of Life The Catalogue of Life is an online database that provides an index of known species of animals, plants, fungi, and microorganisms. It was created in 2001 as a partnership between the global Species 2000 and the American Integrated Taxonomic Info ...
, as a synonym of ''Iris germanica''. It was verified as ''Iris sambucina'' by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
and the
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 9 January 2003, then updated on 12 September 2005.''Iris sambucina'' is a tentatively accepted name by the RHS and listed in the RHS Plant Finder book.


Distribution and habitat

It is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to southern or central Europe.


Range

It is found in Europe, within Germany, GB and Ireland, Italy, Balkans (former Yugoslavia), and Spain.


Habitat

It grows on sunny, well drained rock slopes, in meadows of mountains, and on walls of ancient castles. Within the
South Tyrol it, Provincia Autonoma di Bolzano – Alto Adige lld, Provinzia Autonoma de Balsan/Bulsan – Südtirol , settlement_type = Autonomous province , image_skyline = , image_alt ...
(of Italy), the plants can be found at an altitude of above sea level.


Conservation

It is considered rare, even though it was a wide distribution range.


Cultivation

It is
hardy Hardy may refer to: People * Hardy (surname) * Hardy (given name) * Hardy (singer), American singer-songwriter Places Antarctica * Mount Hardy, Enderby Land * Hardy Cove, Greenwich Island * Hardy Rocks, Biscoe Islands Australia * Hardy, Sout ...
to between
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
Zone 3a (from −39.9 °C (−40 °F)) to Zone 8b (up to −9.4 °C (15 °F)), including Zone 5. It prefers to grow in
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
, well drained, or rich soils, in full sun. It can often been cultivated, and can be grown in mixed flower borders, on the margins of shrubberies or being
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in the garden. It was introduced to Britain in 1658, and was then cultivated in 1748 in the UK, by Mr Philip Miller. Although, it is not often found in bulb nurseries, but a few cultivars can be found in German nurseries. It can though be attacked by
slugs Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...
.


Propagation

It can be propagated by division, due to its hybrid origin, it is
sterile Sterile or sterility may refer to: *Asepsis, a state of being free from biological contaminants * Sterile (archaeology), a sediment deposit which contains no evidence of human activity *Sterilization (microbiology), any process that eliminates or ...
and does not produce seed.


Hybrids and cultivars

It can form hybrids that have flowers in combinations of yellow, red and brown hues. Known cultivars include; Adb-el-Kaker, Abel-Kaver, Cheron, Cherion, Gypsy Queen, Leopold, Leopoldine, Mägdeberg, Nemesis, President Lincoln and Virgil.


Toxicity

Like many other irises, most parts of the plant are poisonous (rhizome and leaves), and if mistakenly ingested can cause stomach pains and vomiting. Handling the plant may cause skin irritation or an allergic reaction.


References


Sources

* Aldén, B., S. Ryman & M. Hjertson Våra kulturväxters namn – ursprung och användning. Formas, Stockholm (Handbook on Swedish cultivated and utility plants, their names and origin). 2009 (Vara kulturvaxt namn) * Czerepanov, S. K. Vascular plants of Russia and adjacent states (the former USSR). 1995 (L USSR) * Huxley, A., ed. The new Royal Horticultural Society dictionary of gardening. 1992 (Dict Gard) * Mathew, B. The Iris. 1981 (Iris) * Tutin, T. G. et al., eds. Flora europaea. 1964–1980 (F Eur) * Walters, S. M. et al., eds. European garden flora. 1986– (Eur Gard F)


External links

{{Taxonbar, from=Q1625281
sambucina ''Sambucina'' is a genus of fungi in the Helotiales order. The relationship of this taxon to other taxa within the order is unknown (''incertae sedis''), and it has not yet been placed with certainty into any family. This is a monotypic In b ...
Flora of Europe Plants described in 1759 Taxa named by Michael Foster (physiologist) Garden plants of Europe