Iris Ruthenica
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''Iris ruthenica'', sometimes called ever blooming iris (in the UK), Russian iris, pilgrim iris and Hungarian iris (in Europe), is a species in the genus ''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants *Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional enti ...
''- subgenus '' Limniris''. It is a
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, with a wide distribution, ranging from eastern Europe to Central Asia. It has grass-like leaves, thick stem and violet or bluish lavender flowers which are marked with violet veining.


Description

''Iris ruthenica'' is very variable and hybrids can look very similar to ''
Iris uniflora ''Iris uniflora'' is a species in the genus ''Iris'', it is also in the subgenus '' Limniris''. It is a rhizomatous perennial, from Russia, Mongolia, China and Korea. It has thin grass-like leaves and stems, and purple, blue-purple or violet flow ...
'', the other species in the ''Iris'' series ''Ruthenicae''. It can be variable with its leaf length and width, and flower height. It has a creeping rhizome,James Cullen, Sabina G. Knees and H. Suzanne Cubey (Editors) (about 3–5 mm in diameter) which is branched and has fibrous roots. The creeping rhizome forms a clump or a grass-like tuft plant. It has bright green leaves, or greyish green leaves. That are tall and thin, and grass-like, measuring between 10 – 40 cm (8–13 in) long and 2 – 6 mm wide. The leaves can grow longer than the flower stem. The plant (stem and flowers) grows to a height of between 3–20 cm (12 in). The thick stem is 2–3 cm wide, can grow to heights of between 3–20 cm. It has the remains of last years leaves at the base of the stem. It blooms in spring, (between May, June and July in the UK), or early to mid summer, with one normally, but occasionally 2 fragrant flowers. The large flowers are between 3–5 cm in diameter, with a cylindrical,
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when ...
tube measuring 0.5–1.5 cm long. The flowers come in a range of blue shades between violet and bluish lavender. Which are marked with violet veining. Like other irises, it has 2 pairs of petals, 3 large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
(outer petals), known as the 'falls' and 3 inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
), known as the 'standards'. The falls (measuring 4.5–5 cm) are white. The standards (measuring 4–6 cm) are almost erect. The bracts (measuring 3–5 cm ) are greenish with pink margins, violet blue stigma, and milky white anthers. It has a globose (globe-like) to ovoid shaped seed capsule (measuring 1.2—1.5 cm) in June–August (after the flowering period is over). Once they are ripe, the seed capsules fully open and all the seeds are dispersed in one movement. Unlike other iris species. The seeds are pyriform (pear-shaped) and have an aril (white appendage on the edge of the seed). The aril disappears soon after and shrivels up.


Biochemistry

As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
, having two sets of
chromosomes A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
. Which can be used to identify hybrids and classification of groupings. It has a chromosome count: 2n=84 (found by Simonet in 1934).


Taxonomy

It is written as 紫苞鸢尾 in Chinese script, and known as 'zi bao yuan wei'. It is named after the region of the '
Ruthenia Ruthenia or , uk, Рутенія, translit=Rutenia or uk, Русь, translit=Rus, label=none, pl, Ruś, be, Рутэнія, Русь, russian: Рутения, Русь is an exonym, originally used in Medieval Latin as one of several terms ...
', in
Transylvania Transylvania ( ro, Ardeal or ; hu, Erdély; german: Siebenbürgen) is a historical and cultural region in Central Europe, encompassing central Romania. To the east and south its natural border is the Carpathian Mountains, and to the west the Ap ...
and
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
, It has several common names; 'ever blooming iris' (in the UK), 'Russian Iris',William Robinson 'pilgrim iris' (sometimes called a synonym of ''Iris ruthenica''),Elizabeth Lawrence and 'Hungarian Iris' in Europe. It is known as ''ungersk iris'' in
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
. ''Iris ruthenica'' was first published by
John Bellenden Ker Gawler John Bellenden Ker, originally John Gawler, was an English botanist born about 1764 in Ramridge, Andover, Hampshire and died in June 1842 in the same town. On 5 November 1804 he changed his name to Ker Bellenden, but continued to sign his name a ...
in Botanical Magazine in 1808. It was later published in 1811, as ''Iris ruthenica'' with the common name 'Pigmy iris' in
Curtis's Botanical Magazine ''The Botanical Magazine; or Flower-Garden Displayed'', is an illustrated publication which began in 1787. The longest running botanical magazine, it is widely referred to by the subsequent name ''Curtis's Botanical Magazine''. Each of the issue ...
, vol. 34, table 1393. Pigmy iris is now used as the common name of ''
Iris pumila ''Iris pumila'', sometimes known as pygmy iris or dwarf iris, probably originated as a natural hybrid between '' Iris pseudopumila'' Boissier & Heldreich and '' I. attica'' Tineo. It ranges from Austria through eastern Europe and the Balkans, in ...
''. It was mentioned the journals of Captain Beechy's Voyage (in 1825),William Jackson Hooker and is mentioned in Cherepanov's Vascular Plants of Russia.Sergeĭ Kirillovich Cherepanov It was verified by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
and the
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 42 October 2014, and is an accepted name by the RHS.


Distribution and habitat

''Iris ruthenica'' is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to a wide region, including temperate Asia and Europe.


Range

It is found in southern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, through Central Asia, (including
Altai Mountains The Altai Mountains (), also spelled Altay Mountains, are a mountain range in Central Asia, Central and East Asia, where Russia, China, Mongolia and Kazakhstan converge, and where the rivers Irtysh and Ob River, Ob have their headwaters. The m ...
and
Turkestan Turkestan, also spelled Turkistan ( fa, ترکستان, Torkestân, lit=Land of the Turks), is a historical region in Central Asia corresponding to the regions of Transoxiana and Xinjiang. Overview Known as Turan to the Persians, western Turke ...
, on the
Tien Shen Tien may refer to: *Tian, also known as Tien or T'ien, the Chinese religious idea of God or heaven *Tian (surname), also romanized as Tien *Tien (TV channel), a Dutch television channel *Tiền, currency used in Vietnam during the 19th and 20th cen ...
mountain range,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
and
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
), to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
. Within Europe, it is found in
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
. It is listed with ''
Iris bloudowii ''Iris bloudowii'' is a species in the genus ''Iris'', it is also in the subgenus of ''Iris'' and in the ''Psammiris'' section. It is a rhizomatous perennial, from Russia, Siberia, Kazakhstan, Mongolia and China, with sickle-shaped leaves, slend ...
'', ''
Iris humilis ''Iris humilis'' is a plant species in the genus ''Iris (plant), Iris'', it is also in the subgenus of ''Iris subg. Iris, Iris'' and in the ''Psammiris'' section. It is a rhizomatous perennial plant, perennial, with a wide distribution range fro ...
'', ''
Iris lactea ''Iris lactea'' is a species in the genus ''Iris'', it is also in the subgenus '' Limniris''. and the series ''Ensatae'', it is the only species in the series. The Japanese water iris, ''Iris ensata'', is actually in series '' Laevigatae''. It i ...
'', ''
Iris sibirica ''Iris sibirica'' ( commonly known as Siberian iris or Siberian flag), is a species in the genus ''Iris''. It is a rhizomatous herbaceous perennial, from Europe (including France, Italy, Switzerland, Austria, Czech Republic, Slovakia, Germany, Hu ...
'', ''
Iris tenuifolia ''Iris tenuifolia'' is a beardless iris in the genus ''Iris'', in the subgenus '' Limniris'' and in the series '' Tenuifoliae'' of the genus. It is a rhizomatous herbaceous perennial, from a wide region over central Asia, including Afghanistan, P ...
'' and ''
Iris tigridia ''Iris tigridia'' is a plant species in the genus ''Iris''; it is also in the subgenus ''Iris'' and in the section ''Pseudoregelia''. It is a rhizomatous perennial, from Kazakhstan, Russia, Mongolia and China. It has dark green or greyish green, ...
'' as being found in the
Altai-Sayan region The Altai-Sayan region is an area of Inner Asia proximate to the Altai Mountains and the Sayan Mountains, near to where Russia, China, Mongolia and Kazakhstan come together. This region is one of the world centers of temperate plant diversity. Its ...
(where Russia, China, Mongolia and Kazakhstan come together).


Habitat

It is found on dry meadows (including grass plains and steppes), pine and birch forest edges and edges of woodland. It can also be found in forest clearings in the forest-meadow mountain belt. Forming a thicket ground-cover. In Mongolia it is found under
Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
/
Betula platyphylla ''Betula platyphylla'', the Asian white birch or Japanese white birch, is a tree species in the family Betulaceae. It can be found in subarctic and temperate Asia in Japan, China, Korea, Mongolia, and Russian Far East and Siberia Siberia ( ...
subtaiga forests, in montane meadow steppes with '' Festuca lenensis'' and '' Artemisia sericea'' and in ''
Pinus sibirica ''Pinus sibirica'', or Siberian pine, in the family Pinaceae is a species of pine tree that occurs in Siberia from 58°E in the Ural Mountains east to 126°E in the Stanovoy Range in southern Sakha Republic, and from Igarka at 68°N in the lower ...
''/''
Picea obovata ''Picea obovata'', the Siberian spruce, is a spruce native to Siberia, from the Ural Mountains east to Magadan Oblast, and from the Arctic tree line south to the Altay Mountains in northwestern Mongolia. Description and uses It is a medium-size ...
'' dark taiga forests (within the upper montane belt with ''
Rubus saxatilis ''Rubus saxatilis'', or stone bramble, is a species of bramble widespread across Europe and Asia from Iceland and Spain east as far as China. It has also been found in Greenland. The green stems are 20–60 cm tall and covered with minute n ...
'' and '' Lathyrus humilis''). Elsewhere in
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
, it is found in the Larch forests of Altai and
Sayan mountains The Sayan Mountains (russian: Саяны ''Sajany''; mn, Соёны нуруу, ''Soyonï nurû''; otk, 𐰚𐰇𐰏𐰢𐰤, Kögmen) are a mountain range in southern Siberia, Russia (Buryatia, Irkutsk Oblast, Krasnoyarsk Krai, Tuva Republic ...
including Tuva). It is also found in
Tuva Tuva (; russian: Тува́) or Tyva ( tyv, Тыва), officially the Republic of Tuva (russian: Респу́блика Тыва́, r=Respublika Tyva, p=rʲɪˈspublʲɪkə tɨˈva; tyv, Тыва Республика, translit=Tyva Respublika ...
n Forests as a subcanopy woody species. On the Altai Mountains, it is found with other mountain flowers including Siberian Dogs-tooth Violet ('' Erythronium krylovii''), Altai Foxtail Lily (''
Eremurus ''Eremurus'' is a genus of deciduous perennial flowers in the family Asphodelaceae. They are also known as the foxtail lilies or desert candles. They are native to eastern Europe in (Russia and Ukraine), and temperate Asia from Turkey to China. ...
''), a variety of
saxifrage ''Saxifraga'' is the largest genus in the family Saxifragaceae, containing about 465 species of holarctic perennial plants, known as saxifrages or rockfoils. The Latin word ''saxifraga'' means literally "stone-breaker", from Latin ' ("rock" or " ...
s, ''
Aquilegia ''Aquilegia'' (common names: granny's bonnet, columbine) is a genus of about 60–70 species of perennial plants that are found in meadows, woodlands, and at higher altitudes throughout the Northern Hemisphere, known for the spurred petals Pu ...
'', ''
Gentiana grandiflora ''Gentiana'' is a genus of flowering plants belonging to the gentian family (Gentianaceae), the tribe Gentianeae, and the monophyletic subtribe Gentianinae. With about 400 species it is considered a large genus. They are notable for their mostl ...
'', ''
Papaver nudicaule ''Papaver nudicaule'', the Iceland poppy, is a boreal flowering plant. Equivalence with ''Papaver croceum'' has been contested. Native to subpolar regions of Asia and North America, and the mountains of Central Asia as well as temperate China ...
'' and the yellow ''
Iris bloudowii ''Iris bloudowii'' is a species in the genus ''Iris'', it is also in the subgenus of ''Iris'' and in the ''Psammiris'' section. It is a rhizomatous perennial, from Russia, Siberia, Kazakhstan, Mongolia and China, with sickle-shaped leaves, slend ...
''. At altitudes of between 1800 and 3600m.


Cultivation

It is
hardy Hardy may refer to: People * Hardy (surname) * Hardy (given name) * Hardy (singer), American singer-songwriter Places Antarctica * Mount Hardy, Enderby Land * Hardy Cove, Greenwich Island * Hardy Rocks, Biscoe Islands Australia * Hardy, Sout ...
to
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
Zone 2, or Zone 3. ''Iris ruthenica'' does not flower very well in the UK. It is best cultivated in fertile soils that do not dry out. It is best suited for Rock Gardens or at the front of a flower border. Although sinks or troughs could be used. It also grows well on dry peat banks. It is tolerant of semi-shade, but prefers full sun. Unlike many other irises, it can only be moved with success, during the spring and summer when it is in full growth. ''Iris ruthenica'' is grown in several Russian botanical gardens including, Barnaul, Ivanovo, Irkutsk, Kirov, Moscow, Rostov-on-Don, St. Petersburg, Stavropol, Tomsk, Omsk, Novosibirsk and Chita.


Propagation

It can be propagated by division or by seed. The seeds should be sown in the autumn and the rhizomes divided in early spring. The seeds germinate fairly quickly and new plants are easily raised. But the young plants must not dry out. The old and damaged rhizomes should be removed before replanting.


Hybrids and cultivars

''Iris ruthenica'' var. ''nana'' was once thought to be a smaller variety of ''Iris ruthenica''. but this is now considered a synonym. Although, ''Iris ruthenica'' var. ''brevituba'' which has a small perianth tube, and violet flowers, it is also now considered a variant. ''Iris ruthenica'' has the following known cultivars; *'Gamlin Blue', *'Red Form'.


References


Sources

*Aldén, B., S. Ryman & M. Hjertson. 2009. Våra kulturväxters namn – ursprung och användning. Formas, Stockholm (Handbook on Swedish cultivated and utility plants, their names and origin). *Czerepanov, S. K. 1995. Vascular plants of Russia and adjacent states (the former USSR). *Khassanov, F. O. & N. Rakhimova. 2012. Taxonomic revision of the genus Iris L. (Iridaceae Juss.) for the flora of Central Asia. Stapfia 97:175. *Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. *Mathew, B. 1981. The Iris. 83. *Tutin, T. G. et al., eds. 1964–1980. Flora europaea. *Waddick, J. W. & Zhao Yu-tang. 1992. Iris of China.


External links

* * {{Taxonbar, from=Q1594646
ruthenica ''Ruthenica'' is a genus of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the subfamily Clausiliinae of the family Clausiliidae, the door snails, all of which have a clausilium The clausilium is a calcareous anatomical ...
Flora of Central Asia Flora of Eastern Europe Flora of China Flora of Kazakhstan Flora of Mongolia Flora of Russia Garden plants of Asia Garden plants of Europe Plants described in 1901 Flora of Korea Flora of Primorsky Krai Flora of Manchuria Flora of Altai (region)