Iris Graminea
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''Iris graminea'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
belonging to the subgenus '' Limniris'' of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants *Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional enti ...
'', in particular the series '' Spuriae''. It is a
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, with purple or violet blue flowers almost hidden by narrow, grass-like leaves, and a plum scented fragrance. It is cultivated as an ornamental plant in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
regions. It has several common names, including grass-leaved flag, grass leaved iris, plum iris and plum tart iris (due to its scent). This species naturally occurs in the southern half of Europe, from
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
and
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
in the West to
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
and the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
in the East.


Description

''Iris graminea'' has slender, short, hard rhizomes.British Iris Society (1997) The plant has narrow grass-like foliage. and can grow up to long and 0.5–1.5 cm wide.Stephen Lacey They can have many branches creating dense tufts, clumps or tussocks of plants. The flower has bright purple flowers, and shiny, green leaves. Some plants can have dark green leaves. They have strongly, visible longitudinal veins. The leaves are longer than the flowering stems.Margery Fish After the plant has flowered, the foliage has the tendency to extend. Later, the
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
foliage dies back during the winter. ''Iris graminea'' has a flattened stem that grows up to long. The stems have 1–2
spathe In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s (leaves of the flower bud), which are unequal in size. The lower is larger and rather leaf-like. The stems hold 1–2 terminal (top of stem) flowers, blooming in spring and summer, between May and June. The stems are normally, unbranched, but (if they have a second flower), the
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
, is up to 6 cm long. It has 2 pairs of petals, 3 large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
(outer petals), known as the 'falls' and 3 inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
, known as the 'standards'. The falls have a long haft or claw, (section closest to the stem) and a small rounded or oval blade. They are long.James Cullen, Sabina G. Knees and H. Suzanne Cubey (Editors) The centre of the blade has a pale yellow, or white central area, which is veined with violet, purple, or blue. Some references describe a dark purple area with white veining. The claw is sometimes winged, and tinged with green or brown, or veined deep reddish-purple. The erect, standards are purple blue, or purple, or red-violet, They are long, with green or brown shade at the base. It has a long and arching, purple style branch, (that is long as the haft), and has a dark purple centre stripe. They have a brown base. It has an ovary with double ridges, a 2 pointed stigma and pollen that is orange-red. It also has a short perianth tube. After the iris has flowered, it produces a long seed capsule, which has a narrowed point. Inside the capsule, are pear shaped seeds, which are slightly compressed and flattened. The flower has been described as having a fruity scent.


Genetics

As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
, having two sets of
chromosomes A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
. This can be used to identify hybrids and classification of groupings. It has been counted several times including, 2n=34, Simonet 1932; 2n=34, Lenz 1963; 2n=34, Popova, M., & I. Cesmedziev, (1975 & 1976); 2n=34, Colasante & Sauer, 1993; 2n=28, 34, 36, Lovka, 1995 and 2n=34, Dobeš et al., 1997. It is generally published as 2n=34.Andreĭ Aleksandrovich Fedoro (Editor)


Taxonomy

It is known as ''Iris à feuilles de graminées'' or ''Iris de Bayonne'' in
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, and ''Giaggiolo susino'' in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
. It is known as ''kosaciec trawolistny'' in
Polish Polish may refer to: * Anything from or related to Poland, a country in Europe * Polish language * Poles, people from Poland or of Polish descent * Polish chicken *Polish brothers (Mark Polish and Michael Polish, born 1970), American twin screenwr ...
, and ''Iris trávolistý'' in Czech and Slovak. and it is known as ''zlakovidny iris'' in Russia, and ''gräsiris'' in
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
,(meaning grass iris). The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''graminea'' refers to ''grassy'' due to the grass-like leaves. It has common names of ''Grass-Leaved Flag'' (in the US) or ''Grass leaved Iris'', ''Plum Iris'', and ''Plum tart Iris'', or ''Plum scented Iris'', (because of the scent). It was originally published and described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' Vol.1 page39 on 1 May 1753. It was later published by Ker-Gawler in Curtis's Botanical Magazine Vol.18 page681 in 1803. Then by B. Fedtsch in ''Flora of SSSR'' Vol.4 page529 in 1935. It was verified by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 4 April 2003 and then updated on 1 December 2004. It is an accepted name by the RHS, and it gained the RHS's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


Distribution

It is native to temperate areas of Europe, and Asia. It is widespread from Spain to Russia. Within Asia, it is found in the Caucasus regions, of Georgia, and the
Russian Federation Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
, (in
Ciscaucasia The North Caucasus, ( ady, Темыр Къафкъас, Temır Qafqas; kbd, Ишхъэрэ Къаукъаз, İṩxhərə Qauqaz; ce, Къилбаседа Кавказ, Q̇ilbaseda Kavkaz; , os, Цӕгат Кавказ, Cægat Kavkaz, inh, ...
and
Dagestan Dagestan ( ; rus, Дагеста́н, , dəɡʲɪˈstan, links=yes), officially the Republic of Dagestan (russian: Респу́блика Дагеста́н, Respúblika Dagestán, links=no), is a republic of Russia situated in the North C ...
). Also in
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
(in Asia Minor). Within Europe, it is found in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
, the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
and
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
, (in
Moravia Moravia ( , also , ; cs, Morava ; german: link=yes, Mähren ; pl, Morawy ; szl, Morawa; la, Moravia) is a historical region in the east of the Czech Republic and one of three historical Czech lands, with Bohemia and Czech Silesia. The me ...
), ex-Yugoslavia, France, Germany, Hungary, Italy, Poland, Romania, Spain, Switzerland, and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
, (in
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a pop ...
,). It may be
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in the Moravia region of the Czech Republic, and in Germany.


Habitat

It is found growing in
scrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, Herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally or ...
, in grass lands, in meadows (or pastures), in open woods, and in rocky or gravelly soils of the mountains.


Conservation

''I. graminea'' is an 'endangered species' in the Czech Republic and Slovakia, it is also listed as 'vulnerable' in Hungary. Within
Saxony Saxony (german: Sachsen ; Upper Saxon: ''Saggsn''; hsb, Sakska), officially the Free State of Saxony (german: Freistaat Sachsen, links=no ; Upper Saxon: ''Freischdaad Saggsn''; hsb, Swobodny stat Sakska, links=no), is a landlocked state of ...
(Germany), it is listed as rare. ''I. graminea'' is thought to be
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
in Poland, but was formerly found near Cieszyn. It has been listed in various
Red Data Books The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
.


Cultivation

''I. graminea'' is
hardy Hardy may refer to: People * Hardy (surname) * Hardy (given name) * Hardy (singer), American singer-songwriter Places Antarctica * Mount Hardy, Enderby Land * Hardy Cove, Greenwich Island * Hardy Rocks, Biscoe Islands Australia * Hardy, Sout ...
to between
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
Zones 3 and 9. It is hardy to Europe Zone H2. It can survive temperatures as low as −20 °C. It is hardy in most places of the UK. During the winter, the foliage dies back, leaving the rhizome under the ground. In very cold climates, a mulch may help the plant survive colder temperatures. It can be grown in normal garden conditions.Benjamin Yoe Morrison They prefer neutral or slightly acidic, (or lime-free), well drained, and fertile (or humus rich) soils. They can tolerate dry soils, or clay and heavy soils, but it prefers damp soils. They prefer positions in full sun, and can tolerate partial shade. It requires adequate watering during growth, such as a moist spring and then drier during the summer. Mature plants can tolerate drought periods. It can be fertilized in early spring and after flowers have bloomed. They can be grown in a front of a mixed border, or rockery. As well as being
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in the garden, they can be also grown in containers, as long as they are well watered. The flowers can be used in bouquets, so can also be grown in a cutting garden. They do not like root disturbance, similar to other spuria irises. It suffers from no serious insect or disease problems.
Crown rot A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
is an infrequently occurring disease problem. It is susceptible to certain viruses, such as bacterial leaf blight, soft rot, rhizome rot,
leaf spot A leaf spot is a limited, discoloured, diseased area of a leaf that is caused by fungal, bacterial or viral plant diseases, or by injuries from nematodes, insects, environmental factors, toxicity or herbicides. These discoloured spots or lesions ...
,
rust Rust is an iron oxide, a usually reddish-brown oxide formed by the reaction of iron and oxygen in the catalytic presence of water or air moisture. Rust consists of hydrous iron(III) oxides (Fe2O3·nH2O) and iron(III) oxide-hydroxide (FeO(OH ...
, viruses and
scorch Scorch may refer to: Entertainment * Scorch, a dragon puppet character created by Ronn Lucas ** ''Scorch'' (TV series), a 1992 CBS sitcom starring the Ronn Lucas character * Scorch (comics), a.k.a. Aubrey Sparks, an antagonist of Superman * Sco ...
. It is also can be susceptible to damage by
Iris borer Iris most often refers to: *Iris (anatomy), part of the eye * Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants *Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
, verbina bud moth, white flies, iris weevil,
thrips Thrips ( order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Different thrips species feed mostly on plants by puncturing and sucking up the contents, although a few are ...
,
slugs Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...
,
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
s, aphids and
nematode The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhab ...
s. It can also tolerate damage from deer. Aphid ''
Aphis newtoni ''Aphis'' is a genus of insects in the family Aphididae containing at least 600 species of aphids. It includes many notorious agricultural pests, such as the soybean aphid '' Aphis glycines''. Many species of ''Aphis'', such as '' A. coreopsidis ...
'' can be found on the plant.R. L. Blackman, Victor F. Eastop The iris has been cultivated since at least 1568. It was first grown in
Cambridge Botanic Garden The Cambridge University Botanic Garden is a botanical garden located in Cambridge, England, associated with the university Department of Plant Sciences (formerly Botany School). It lies between Trumpington Road to the west, Bateman Street to ...
in 1733, where it was labelled as ''narrow leafed plum scent iris''. For many years it was grown in St. Petersburg Botanical Garden, and the botanical garden of
Ufa Ufa ( ba, Өфө , Öfö; russian: Уфа́, r=Ufá, p=ʊˈfa) is the largest city and capital of Bashkortostan, Russia. The city lies at the confluence of the Belaya and Ufa rivers, in the centre-north of Bashkortostan, on hills forming the ...
. It is the most commonly grown ''Iris spuria'' species, and can be easily found in plants nurseries, though it is most commonly known in Europe.


Propagation

It can also be propagated by division or by seed growing. It rarely needs lifting and dividing. Large clumps of mature plants can be lifted between mid-summer to early autumn, then divided before being replanted. As it does not like root disturbance, so seed growing is preferred. If re-planted, they can take a year to 'settle' before beginning to bloom again. Seeds are collected from the pods after flowering. Seeds are then sown in containers in a cold frame in autumn, or between September and May. Seeds should be kept moist for about 2–4 weeks and also warm (at about 15–20 °C). Then temperatures between −4 °C and + 4 °C suspend for 4–6 weeks. These conditions are best reached at the autumn sowing outdoors. Seed trays should then be kept at temperatures of 5 °C to + 10 °C after the cooling period, for a few weeks. In the wild, these freezing and heat period) conditions take place naturally. The plants need to grow and mature for about 2–3 years before they begin to flower.


Hybrids and cultivars

''Iris graminea'' has the following known varieties: 'Achtaroffii', 'Adami', 'Colchica', 'Graminea Lamprophylla', 'Graminea Sylvatica', 'Graminea latifolia', 'Gravenia', 'Hort's variety', and 'Pseudocyperus'. The latter of these has ranges from the Czech Republic, Romania, and Slovakia. ''Pseudocyperus'' has broad leaves, but no scent.


References


Sources

* * ** * * * **Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. **Tutin, T. G. et al., eds. 1964–1980. Flora Europaea.


External links


BioLib

Iris graminea
{{Taxonbar, from=Q161412 graminea Flora of Europe Flora of the Caucasus Garden plants of Europe Plants described in 1753 Taxa named by Carl Linnaeus