Iris Cengialti
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''Iris pallida'' subsp. ''cengialti'' is a subspecies in the genus ''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants *Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional enti ...
'', it is also in the subgenus ''
Iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants *Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional enti ...
''. It is a
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
, from
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
and (part of the former country of
Yugoslavia Yugoslavia (; sh-Latn-Cyrl, separator=" / ", Jugoslavija, Југославија ; sl, Jugoslavija ; mk, Југославија ;; rup, Iugoslavia; hu, Jugoszlávia; rue, label=Pannonian Rusyn, Югославия, translit=Juhoslavija ...
)
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
. It has yellowish-green, glaucous, lanceolate or ensiform leaves, tall stem, green flushed with purple spathes, 2 short branches, 2–3 scented flowers, in shades of violet, blue-violet, deep purple, blue-purple, deep blue-purple, pale purple, deep blue, to mid-blue. It has a yellow or orange tipped beard. It was originally published as ''Iris cengialti'' but then re-classified as a subspecies of ''
Iris pallida ''Iris pallida'', the Dalmatian iris or sweet iris, is a hardy flowering perennial plant of the genus '' Iris'', family Iridaceae. It is native to the Dalmatian coast (Croatia) but widely naturalised elsewhere. It is a member of the subgenus '' I ...
'', and known as ''Iris pallida'' subsp. ''cengialti'', but it is often still called ''Iris cengialti''. It is cultivated as an ornamental garden plant in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
regions.


Description

It has a long, stout,Richard Lynch fleshy, light-coloured (underground) rhizome. That is 1–3 cm wide (in diameter), and has long secondary roots. It forms creeping plants. It has yellowish-green, lanceolate, or ensiform (sword-shaped), leaves, that are glaucous. The basal leaves, can grow up to between long, and about 1.3 cm wide. The herbaceous leaves (die in the winter), sheath the stem. It has a round (in section) stem, or peduncle, that can grow up to between , or tall. The stems are taller than the leaves, and at higher levels on the mountains, the plants are shorter. It is also shorter than ''Iris pallida''. The stem has glaucous green and ensiform spathes (leaves of the flower bud). They are slightly flushed with purple, and before flowering, they become pale brown, (scarious) membranous, and papery, They are 2.5 cm long, and between wide. It has 2 short branches (or
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
s). The stems (and the branches) hold between 2 and 3 flowers, It can have up to 6 flowers, but normally has 3 flowers, in spring, between April to June, or May, to July. The small, scented flowers, are in diameter, come in shades of blue, from violet, blue-violet, deep purple, blue-purple, deep blue-purple, pale purple, deep blue, to mid-blue. Like other irises, it has 2 pairs of petals, 3 large
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
(outer petals), known as the 'falls' and 3 inner, smaller petals (or
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
), known as the 'standards'. The falls are obovate or cuneate shaped, they are long and wide. In the centre of the falls, is a short, (16–30 mm,) thick row of small hairs (the beard), which is yellowish-white, bright yellow, or white with orange tips. The standards are a similar size to the falls. It has 2.5 cm long style branch, that is paler than the falls and standards, and has deltoid crests. It also has a six grooved, ellipsoid ovary, and a 0.6–0.8 cm long perianth tube. After the iris has flowered, it produces an ovoid seed capsule, that is long and 1 cm wide, with 3 sections, which contain 15–20 oval grey or yellowish seeds.


Genetics

In 1956, a
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
analysis was carried out on 40 species of Iris, belonging to the subgenera ''Eupogoniris'' and ''Pogoniris''. It found that 24-chromosome tall bearded species could be divided into 3 karyotypes of ''Iris pallida''. ''
Iris kashmiriana ''Iris kashmiriana'' is a plant species in the genus '' Iris'', it is also in the subgenus '' Iris''. It is a rhizomatous perennial, from Kashmir, India. It has straight, sword-shaped, glaucous leaves, tall, thick stem with up 2 short branches, ...
'' has 2 pairs of median-constricted marker chromosomes, ''
Iris illyrica Iris illyrica, the Illyrian iris, is a perennial plant from the iris family (Iridaceae), native to Southeastern Europe. Distribution Its native range consists of much of the ancient region Illyria, for which it is named, located on the Balkan Pe ...
'', ''Iris cengialti'', and ''
Iris imbricata ''Iris imbricata'' is a species in the genus '' Iris'', it is also in the subgenus '' Iris''. It is a rhizomatous perennial, from the Caucasus mountains, within Iran, Armenia, Azerbaijan and Georgia. It has broad, sword-like, yellow green or li ...
'', lastly ''
Iris variegata ''Iris variegata'', commonly known as the Hungarian iris, is a plant species in the genus ''Iris'', also in the subgenus ''Iris''. It is a rhizomatous perennial from eastern Europe. It has dark green, ribbed leaves. The branched flowering stems c ...
'', ''Iris reginae'' (later classified as a synonym of ''Iris variegata''), and ''
Iris perrieri ''Iris perrieri'' is a plant species in the genus '' Iris''; it is also in the subgenus '' Iris''. It is a rhizomatous perennial, from the Savoy Alps in southern France and recently in Italy. It has green, deeply ribbed, sickle shaped leaves, a ...
'' all have no median-constricted chromosomes. As most irises are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
, having two sets of
chromosomes A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
, this can be used to identify hybrids and classification of groupings. It has a chromosome count: 2n=24.


Taxonomy

It is commonly known as 'Iris del Cengio', 'Monte Cengio iris'. In Italy, it is known as 'Giaggiolo del Cengio Alto'. In
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
, it is known as 'Bohinj Iris', or 'Bòcje of lof'. In
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
, it is known as 'Rovereto Schwertlilie'. It is sometimes misspelt as ''Iris cengualti''. The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''cengialti'' refers to a mountain or large hill 'Mount Cengialto' (also known as Monte 'Cengio Alto',), close to the town of
Rovereto Rovereto (; "wood of sessile oaks"; locally: ''Roveredo'') is a city and ''comune'' in Trentino in northern Italy, located in the Vallagarina valley of the Adige River. History Rovereto was an ancient fortress town standing at the frontier b ...
, in northern Italy. Confusingly, another mountain near
Arsiero Arsiero is a town in the province of Vicenza, Veneto, Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its te ...
in the
province of Vicenza The Province of Vicenza ( it, Provincia di Vicenza) is a province in the Veneto region in northern Italy. Its capital city is Vicenza. The province has an area of 2,722.53 km², and a total population of 865,082 (as of 2017). There are 1 ...
(Veneto), is also known as Monte Cengio Alto. It was found by
Francesco Ambrosi Francesco Ambrosi (November 17, 1821 – 9 April 1897) was an Italian botanist, librarian, ethnologist and historian. Biography Ambrosi was born to a family of herders in Borgo Valsugana, the present day Italian province of Trentinoat that tim ...
in 1854, on Monte Cengialto, It was then published as ''Iris cengialti'' by Ambrosi in Flora del Tirolo Meridionale Vol.1 (Fl. Tirolo Mer.) on page 643 in 1854, then fully published and described by A. Kerner in Österreichische Botanische Zeitschrift (Oesterr. Bot. Z.) Vol. 21, No.9, on pages 225–231 in September 1871. A. Kerner was then assigned the author of the name ''Iris cengialti'', previously ascribed to Ambrosi. ''Iris pallida'', ''Iris cengialti'' and ''
Iris illyrica Iris illyrica, the Illyrian iris, is a perennial plant from the iris family (Iridaceae), native to Southeastern Europe. Distribution Its native range consists of much of the ancient region Illyria, for which it is named, located on the Balkan Pe ...
'' are all considered closely related but often given different taxonomic rank (by various authors Kerner, 1871; Ambrosi, 1854; Foster, 1886; Tommasini, 1875; Pampanini, 1909; Lausi, 1964; Mathew, 1981; Colasante, 1995: Terpin et al., 1996). In May 1886, Sir Michael Foster noted in
Gardeners' Chronicle ''The Gardeners' Chronicle'' was a British horticulture periodical. It lasted as a title in its own right for nearly 150 years and is still extant as part of the magazine ''Horticulture Week''. History Founded in 1841 by the horticulturists Josep ...
on pages554 and 555, that it had similarities with ''Iris pallida''.
William Rickatson Dykes William Rickatson Dykes (4 November 1877 – 1 December 1925) was an English amateur botanist who became an expert in the field of iris breeding and wrote several influential books on the subject. He was also interested in tulips, amaryllis, and ...
in his book in 'Handbook of Garden Irises' in 1914, suggested that ''Iris cengialti'' and ''Iris pallida'' were connected and should be merged under ''Iris pallida''. In 1939,
Brian Mathew Brian Frederick Mathew MBE, VMH is a British botanist, born in the village of Limpsfield, Surrey, England. His particular area of expertise is bulbous plants, particularly ornamental bulbous plants, although he has contributed to other fields o ...
considered it to be a subspecies of ''Iris pallida''. Then in Giorn. Bot. Ital. Vol.130 on page575 in 1996, ''Iris cengialti'' was classified as a synonym of ''Iris pallida'' subsp. ''cengialti''. It is normally classified as ''Iris pallida'' subsp. ''cengialti'', and thought to be a naturally occurring hybrid. In Slovenia, a form of the plant is called ''Iris cengialti vochinensis'', or ''Iris pallida'' subsp. ''cengialti'' f. ''vochinensis''. probably of Asia Minor origin, It was verified as ''Iris pallida'' subsp. ''cengialti'' by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
and the
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 27 February 2002, and then updated on 1 December 2004. It is listed in the
Encyclopedia of Life The ''Encyclopedia of Life'' (''EOL'') is a free, online encyclopedia intended to document all of the 1.9 million living species known to science. It is compiled from existing trusted databases curated by experts and with the assistance of non ...
, and listed on the
Catalogue of Life The Catalogue of Life is an online database that provides an index of known species of animals, plants, fungi, and microorganisms. It was created in 2001 as a partnership between the global Species 2000 and the American Integrated Taxonomic Info ...
as ''Iris pallida'' subsp. ''cengialti''.


Distribution and habitat

It is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to Europe.


Range

It endemic in the alpine region, on the
Dolomites The Dolomites ( it, Dolomiti ; Ladin: ''Dolomites''; german: Dolomiten ; vec, Dołomiti : fur, Dolomitis), also known as the Dolomite Mountains, Dolomite Alps or Dolomitic Alps, are a mountain range located in northeastern Italy. They form par ...
(mountain range).Erika Pignatti and Sandro Pignatti It is found in north east Italy, (within the
region In geography, regions, otherwise referred to as zones, lands or territories, are areas that are broadly divided by physical characteristics (physical geography), human impact characteristics (human geography), and the interaction of humanity and t ...
of
Trentino Trentino ( lld, Trentin), officially the Autonomous Province of Trento, is an autonomous province of Italy, in the country's far north. The Trentino and South Tyrol constitute the region of Trentino-Alto Adige/Südtirol, an autonomous region ...
, and
Veneto Veneto (, ; vec, Vèneto ) or Venetia is one of the 20 regions of Italy. Its population is about five million, ranking fourth in Italy. The region's capital is Venice while the biggest city is Verona. Veneto was part of the Roman Empire unt ...
) and (part of the former Yugoslavia), in Slovenia, (within the regions of
Bohinj Bohinj (; german: Wochein), or the Bohinj Valley ( sl, Bohinjska dolina) or Bohinj Basin ( sl, Bohinjska kotlina), is a 20 km long and 5 km wide basin in the Julian Alps, in the Upper Carniola region of northwestern Slovenia. It is trav ...
. and
Istria Istria ( ; Croatian language, Croatian and Slovene language, Slovene: ; ist, Eîstria; Istro-Romanian language, Istro-Romanian, Italian language, Italian and Venetian language, Venetian: ; formerly in Latin and in Ancient Greek) is the larges ...
). In Trentino, it has found on
Monte Zugna Coni Zugna, also known as Monte Zugna, is a mountain in the Vicentine Alps, in northeastern Italy. It has an elevation of 1,865 metres and is located near the southern border of the province of Trento, close to the province of Vicenza, just nort ...
(south of Rovereto), and Mount Brione. In Veneto, it can be found on Mount Summano, in the
province of Belluno The Province of Belluno ( it, Provincia di Belluno; de-AT, Provinz Belluno; lld, Provinzia de Belum) is a province in the Veneto region of Italy. Its capital is the city of Belluno. It has an area of and a total population of about 205,000. ...
.


Habitat

It grows on the rocky mountain sides, on
scree Scree is a collection of broken rock fragments at the base of a cliff or other steep rocky mass that has accumulated through periodic rockfall. Landforms associated with these materials are often called talus deposits. Talus deposits typically ha ...
, in meadows, and dry grasslands. Usually on
karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
, or limestone soils. They can be found at an altitude of above sea level. They can found in the Dolomites, with other plants including; '' alyssum ovirense'', '' androsace villosa'', '' centaurea haynaldii'', '' crepis froelichiana subsp. dinarica'', ''
eritrichium nanum ''Eritrichium nanum'', the arctic alpine forget-me-not or king-of-the-Alps, is a circumpolar alpine cushion plant which occurs in the North American Rocky mountains as well as the European Alps. It grows at elevations of 10,000 feet in an environ ...
'', ''
euphorbia kerneri ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
'', ''
geranium argenteum ''Geranium argenteum'', the silvery crane's bill, is an ornamental plant in the family Geraniaceae, which is native to Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical ...
'', ''
leontodon berinii ''Leontodon'' is a genus of plants in the tribe Cichorieae within the family Asteraceae, commonly known as hawkbits. Their English name derives from the mediaeval belief that hawks ate the plant to improve their eyesight. Although originally o ...
'', ''
lilium carniolicum ''Lilium carniolicum'', commonly called golden apple or Carniolan lily, is a lily native to the Balkans, as well as to Austria and northeastern Italy (Veneto + Friuli Venezia Giulia). The species is named after the historical region of Carniola, ...
'', '' potentilla incana'' and ''
thlaspi minimum ''Thlaspi'', or pennycress, is a genus of herbs of temperate regions of the Eurasian continent. They occur in Central and South Europe, South-West Asia and two species are endemic to China. The ''Thlaspi'' has been proven to be a hyperaccumulato ...
''. They also grow in the Belluno forests of ''
Acer pseudoplatanus ''Acer pseudoplatanus'', known as the sycamore in the British Isles and as the sycamore maple in the United States, is a species of flowering plant in the Sapindus, soapberry and lychee family Sapindaceae. It is a large deciduous, broad-leaved ...
'' (Mount Maple) and ''
Tilia platyphyllos ''Tilia platyphyllos'', the large-leaved lime or large-leaved linden, is a species of flowering plant in the family Malvaceae (Tiliaceae). It is a deciduous tree, native to much of Europe, including locally in southwestern Great Britain, growing ...
'' (Lime tree) with other geophytes and ''
Leontodon tenuiflorus ''Leontodon'' is a genus of plants in the tribe Cichorieae within the family Asteraceae, commonly known as hawkbits. Their English name derives from the mediaeval belief that hawks ate the plant to improve their eyesight. Although originally on ...
'' (Dandelion insubrico), ''
Knautia persicina ''Knautia'' is a genus of flowering plants in the family Caprifoliaceae. The common names are variants of "widow flower". Others are given the name "scabious", which properly belongs to the related genus (''Scabiosa''). The name ''Knautia'' come ...
'', ''
Globularia nudicaulis ''Globularia'' is a genus of about 22 species of flowering plants in the family Plantaginaceae, native to central and southern Europe, Macaronesia, northwest Africa and southwest Asia. They are dense low evergreen mat-forming perennials or subs ...
'' and ''
Paeonia officinalis ''Paeonia officinalis'', the common peony, or garden peony, is a species of flowering plant in the family Paeoniaceae, native to mainly mountainous areas of Southern Europe and introduced in Central and Western Europe and North America. ''Paeon ...
'' (Wild Peony). In Slovenia, in Bohinj, it grows in meadows with another rare plant, ''
Linnaea borealis ''Linnaea borealis'' is a species of flowering plant in the family Caprifoliaceae (the honeysuckle family). Until 2013, it was the only species in the genus ''Linnaea''. It is a boreal to subarctic woodland subshrub, commonly known as twinflower ...
'' (twinflower).


Conservation

It is a rare plant, although locally it is abundant on the mountains, growing in masses, hanging onto relatively steep slopes. In Italy, it is listed as a protected species of regional interest (Annex B, LR 9/2007, Art. 96), and would be classed as 'vulnerable' on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
.


Cultivation

It is hardy to 0 °C. It prefers could grown in rich (in fertile loan,), well drained soil in full sun. It can be grown the rock garden. It is generally not attacked by pests and diseases. A herbarium specimen can be found at the
Museum National d'Histoire Naturelle The French National Museum of Natural History, known in French as the ' (abbreviation MNHN), is the national natural history museum of France and a ' of higher education part of Sorbonne Universities. The main museum, with four galleries, is loc ...
in Paris.


Propagation

Irises can generally be propagated by
division Division or divider may refer to: Mathematics *Division (mathematics), the inverse of multiplication *Division algorithm, a method for computing the result of mathematical division Military *Division (military), a formation typically consisting ...
. They should be lifted every three or four years, after the flowers are over. The tuber can be divided, with the dead roots trimmed and removed. The remains can then be replanted in enriched soil (with added compost). The leaves should be cut back to half their length, to prevent wind rock.


Hybrids and cultivars

It has been used in hybridizing (or breeding programmes), and the hybrids that have ''Iris cengialti'' as a parent, can have multiple flowers, on average between 5 and 8. The branches are generally short on slender stems.Benjamin Yoe Morrison


Toxicity

Like many other irises, most parts of the plant are poisonous (rhizome and leaves), if mistakenly ingested can cause stomach pains and vomiting. Also handling the plant may cause a skin irritation or an allergic reaction.David G Spoerke and Susan C. Smolinske


References


Sources

* Mathew, B. 1981. The Iris. 24. ists as ''Iris cengialtii'' Ambrosi * Pignatti, S. 1982. Flora d'Italia. ccepts subsp.* Terpin, K. et al. 1996. Author and type of the name ''Iris cengialti'' (Iridaceae). Giorn. Bot. Ital. 130:575–578. * Tutin, T. G. et al., eds. 1964–1980. Flora europaea. ists as ''I. cengialti'' Ambrosi


External links


Has a large image of the flower in the Belluno Dolomites National Park in Italy

Painting of the iris, from the collection of Botanical Garden – University of Padova
* * {{Taxonbar, from=Q22285048 pallidula subsp. cengialti Plants described in 1871 Garden plants Flora of Europe Flora of Italy Flora of Slovenia Plant subspecies