In biology, an integument is the tissue surrounding an organism's body or an organ within, such as

skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...

, a husk, shell, germ or

rind Rind may refer to: Food *Peel (fruit), or outer covering of any vegetable *Pork rind *The outer layer of cheese *Candied rind; see Succade *Grated rind; see Zest (ingredient) Other uses *Rind (Baloch tribe), a tribe in Pakistan *Rind (giantess), ...

Etymology

The term is derived from ''integumentum'', which is

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...

for "a covering". In a transferred, or figurative sense, it could mean a

cloak A cloak is a type of loose garment worn over clothing, mostly but not always as outerwear for outdoor wear, serving the same purpose as an overcoat, protecting the wearer from the weather. It may form part of a uniform. Cloaks have been and ...

or a disguise. In English, "integument" is a fairly modern word, its origin having been traced back to the early seventeenth century; and refers to a material or layer with which anything is enclosed, clothed, or covered in the sense of "clad" or "coated", as with a skin or husk.

Botanical usage

botany Botany, also called plant science (or plant sciences), plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "bot ...

, the term "integument" may be used as it is in zoology, referring to the covering of an organ. When the context indicates nothing to the contrary, the word commonly refers to an envelope covering the nucellus of the

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the f ...

. The integument may consist of one layer (unitegmic) or two layers (bitegmic), each of which consisting of two or more layers of cells. The integument is perforated by a pore, the micropyle, through which the pollen tube can enter. It may develop into the testa, or

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...

coat.

Zoological usage

The integument of an organ in

zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...

typically would comprise membranes of connective tissue such as those around a kidney or liver. In referring to the integument of an animal, the usual sense is its skin and its derivatives: the

integumentary system The integumentary system is the set of organs forming the outermost layer of an animal's body. It comprises the skin and its appendages, which act as a physical barrier between the external environment and the internal environment that it serves ...

, where "integumentary" is a synonym of "

cutaneous Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other animal coverings, such as the arthropod exoskeleton, have different ...

". In

arthropods Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chitin, oft ...

, the integument, or external "skin", consists of a single layer of epithelial ectoderm from which arises the cuticle, an outer covering of

chitin Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chit ...

, the rigidity of which varies as per its chemical composition.

Derivative terms and sundry usages

Derivative terms include various adjectival forms such as integumentary (e.g. system), integumental (e.g. integumental glands, "peltate glands, the integument being raised like a bladder due to abundant secretion"Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928) and integumented (as opposed to bare). Other illustrative examples of usage occur in the following articles: * Connective tissue in skeletal muscle * Dorsal artery of the penis (example of integument enclosing an internal organ) * Flensing *

Flesh Flesh is any aggregation of soft tissues of an organism. Various multicellular organisms have soft tissues that may be called "flesh". In mammals, including humans, ''flesh'' encompasses muscles, fats and other loose connective tissues, but ...

(generic use of plural "integuments") * Herzog & de Meuron (figurative usage) *

Integumentary system The integumentary system is the set of organs forming the outermost layer of an animal's body. It comprises the skin and its appendages, which act as a physical barrier between the external environment and the internal environment that it serves ...

(senses related to vertebrate skin and medical aspects) * Integumental muscles *

Mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Ma ...

Seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...

References

{{Authority control Animal anatomy Skin Organs (anatomy) Plant anatomy