''Ilex ambigua'' is a species of flowering plant in the holly family known by the common names Carolina holly and sand holly. It is native to the southeastern and south-central United States, along the

coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Coa ...

from

North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

, inland as far as

Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...

Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...

, and

Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...

.Sullivan, Janet. 1994
''Ilex ambigua''.
In: Fire Effects Information System, nline United States Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory.Biota of North America Program 2014 county distribution map
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Description

''Ilex ambigua'' is a large

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

or small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

up to tall. The branches are covered in shiny dark brown or black bark which becomes flaky with age. The twigs are purple.''Ilex ambigua''.
The Nature Conservancy. Some branches have a thick coat of fine hairs. The leaves are up to 18 centimeters (7.2 inches) long by 7 cm (2.8 inches) wide. The leaf margins are partially or entirely toothed or wavy. The species is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

, with male and female reproductive parts occurring on separate individuals. The fruit is a spherical red

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

. The seeds are dispersed by animals, which eat the fruits. ;Habitat ''Ilex ambigua'', grows in many types of sandy habitat, such as sand scrub and

hammocks A hammock (from Spanish , borrowed from Taíno and Arawak ) is a sling made of fabric, rope, or netting, suspended between two or more points, used for swinging, sleeping, or resting. It normally consists of one or more cloth panels, or a wov ...

and hardwood forests and woodlands. It may grow with

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

s such as loblolly,

slash Slash may refer to: * Slash (punctuation), the "/" character Arts and entertainment Fictional characters * Slash (Marvel Comics) * Slash (''Teenage Mutant Ninja Turtles'') Music * Harry Slash & The Slashtones, an American rock band * Nash ...

, and

shortleaf pine ''Pinus echinata'', the shortleaf pine, is a species of pine native to the southeastern United States. Description The tree is variable in form, sometimes straight, sometimes crooked, with an irregular crown. The tree reaches heights of with a ...

, and

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

species. It sometimes grows with its relative,

American holly ''Ilex opaca'', the American holly, is a species of holly, native to the eastern and south-central United States, from coastal Massachusetts south to central Florida, and west to southeastern Missouri and eastern Texas. Description ''Ilex opaca ...

. ;forms and varieties *''Ilex ambigua'' f. ''ambigua'' *''Ilex ambigua'' var. ''ambigua'' *''Ilex ambigua'' f. ''mollis'' (A. Gray) H.E. Ahles *''Ilex ambigua'' var. ''monticola'' (A. Gray) Wunderlin & Poppleton

References

{{Taxonbar, from=Q5997640 ambigua Flora of the Southern United States Plants described in 1803