Ignavusaurus
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''Ignavusaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of basal
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived during the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
in what is now
Lesotho Lesotho ( ), officially the Kingdom of Lesotho, is a country landlocked country, landlocked as an Enclave and exclave, enclave in South Africa. It is situated in the Maloti Mountains and contains the Thabana Ntlenyana, highest mountains in Sou ...
. Its fossils were found in the
Upper Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, sout ...
which is probably
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
in age (around 200
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
). It was described on the
basis Basis may refer to: Finance and accounting *Adjusted basis, the net cost of an asset after adjusting for various tax-related items *Basis point, 0.01%, often used in the context of interest rates *Basis trading, a trading strategy consisting of ...
of a partial, well preserved articulated skeleton. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''I. rachelis'', was described in 2010 by Spanish palaeontologist F. Knoll.


Discovery

The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
was discovered in southern
Lesotho Lesotho ( ), officially the Kingdom of Lesotho, is a country landlocked country, landlocked as an Enclave and exclave, enclave in South Africa. It is situated in the Maloti Mountains and contains the Thabana Ntlenyana, highest mountains in Sou ...
near Ha Ralekoala. The fossils were discovered in a mostly articulated state, although the skull was badly damaged, having broken into more than 120 fragments. An unpublished 2002 Ph.D. dissertation referred to the remains, but they were not formally described for several years, and the find remained unpublished. In 2010, the fossils were described by F. Knoll of the
Museo Nacional de Ciencias Naturales The National Museum of Natural Sciences ( es, Museo Nacional de Ciencias Naturales) is the national museum of natural history of Spain. It is situated in the center of Madrid, by the Paseo de la Castellana. It is managed by the Spanish National Re ...
-CSIC in
Madrid Madrid ( , ) is the capital and most populous city of Spain. The city has almost 3.4 million inhabitants and a metropolitan area population of approximately 6.7 million. It is the second-largest city in the European Union (EU), and ...
as a new taxon, ''Ignavusaurus rachelis''. The fossils were brought to the National Museum of Natural History in Paris, where they were provisionally catalogued as BM HR 20; the plan was to return the fossils to Lesotho upon the opening of the Lesotho National Museum.


Etymology

The generic name ''Ignavusaurus'' is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word ''ignavus'' ("coward") and
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''sauros'' ("lizard"). It refers to the type locality – Ha Ralekoala, that literally means "The place of the father of the coward". The specific name of the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''rachelis'', honours Spanish palaeontologist Raquel López-Antoñanzas.


Description

The holotype had a body length of about and weighed about .
Histological Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
analysis of the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
and
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
indicates that BM HR 20 was fast-growing individual, maybe less than one year old, based on age determination studies. Like other early sauropodomorphs, ''Ignavusaurus'' had a long, slender neck and tail. It generally resembled ''
Massospondylus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
'' and ''
Melanorosaurus ''Melanorosaurus'' (meaning "Black Mountain Lizard", from the Greek ''melas/'', "black", ''oros/'', "mountain" + ''/'', "lizard") is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Af ...
'', two other sauropodomorphs from southern Africa, although the shape and position of the teeth, among other factors, led Knoll to conclude that the remains belong to a previously unknown genus.


Classification

''Ignavusaurus'' is a basal
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...
, a member of the long-necked,
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
,
saurischian Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithis ...
dinosaurs. Although originally recovered as more primitive than
Plateosauria Plateosauria is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to the Late Cretaceous. The name Plateosauria was first coined by Gustav Tornier in 1913. The name afterwards fell out of use until the 1980s. Classificat ...
by Knoll (2010), Yates et al. (2011) noted that some characters used to place ''Ignavusaurus'' outside Massopoda are reflective of juvenile status and instead are more similar to juvenile ''Massospondylus'', considering ''Ignavusaurus'' probably synonymous with ''
Massospondylus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
''. Meanwhile, a
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis presented by Apaldetti et al. (2011) found ''Ignavusaurus'' to be a valid genus which is most closely related to ''
Sarahsaurus ''Sarahsaurus'' is a genus of basal sauropodomorph dinosaur which lived during the Early Jurassic period in what is now northeastern Arizona, United States. Discovery and naming All specimens of ''Sarahsaurus'' were collected from the Lower Ju ...
'' within
Massopoda Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, define ...
, somewhat agreeing with Yates et al. that ''Ignavusaurus'' is not as primitive as originally thought. A cladistic analysis conducted by Chapelle and Choiniere (2018) in their paper updating knowledge of the skull of ''Massospondylus'' recovered ''Ignavusaurus'' along with ''Sarahsaurus'' in Massospondylidae, as did the cladistic analysis of Chapelle et al. (2019).Chapelle KEJ, Barrett PM, Botha J, Choiniere JN. 2019. ''Ngwevu intloko'': a new early sauropodomorph dinosaur from the Lower Jurassic Elliot Formation of South Africa and comments on cranial ontogeny in Massospondylus carinatus. PeerJ 7:e7240 https://doi.org/10.7717/peerj.7240


References

{{Taxonbar, from=Q93603 Massopoda Early Jurassic dinosaurs of Africa Fossil taxa described in 2010 Paleontology in Lesotho