''Iberomesornis'' ("Spanish intermediate bird") is a
monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
enantiornithine
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
of the
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
of
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, i ...
.
Discovery
In 1985 the
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
of ''Iberomesornis'' was discovered by
Armando Díaz Romeral in the
Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma.
Geology
Pro ...
Calizas de La Huérguina Formation at
Las Hoyas,
Cuenca Province
Cuenca is one of the five provinces of the autonomous community of Castilla-La Mancha. It is located in the eastern part of this autonomous community and covers 17,141 square km. It has a population of 203,841 inhabitants -- the least populated o ...
, east central Spain, which dates to the late
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
, roughly 125 million years ago. The find was first reported in 1988. In 1992 the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Iberomesornis romerali'' was named and described by
José Luis Sanz and
José Fernando Bonaparte
José is a predominantly Spanish and Portuguese form of the given name Joseph. While spelled alike, this name is pronounced differently in each language: Spanish ; Portuguese (or ).
In French, the name ''José'', pronounced , is an old vernacul ...
. The generic name is derived from
Iberia
The Iberian Peninsula (),
**
* Aragonese and Occitan: ''Peninsula Iberica''
**
**
* french: Péninsule Ibérique
* mwl, Península Eibérica
* eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and
Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
, , "middle", and , , "bird", in reference to the intermediate status between the most basal and the modern birds. The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
honours Romeral.
The
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen, LH-22, part of the Las Hoyas Collection, consists of a compressed articulated partial skeleton of an adult individual lacking the skull, the anterior neck and most of the hands. A second specimen, LH-8200, was referred to a ''Iberomesornis'' sp. in 1994, consisting of the left foot of an individual similar in size to the holotype. After further preparation of the fossil, the species was redescribed by
Paul Sereno
Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...
in 2000.
Biology and ecology
''Iberomesornis'' was quite small, no bigger than a large modern
sparrow
Sparrow may refer to:
Birds
* Old World sparrows, family Passeridae
* New World sparrows, family Passerellidae
* two species in the Passerine family Estrildidae:
** Java sparrow
** Timor sparrow
* Hedge sparrow, also known as the dunnock or hedg ...
. On the assumption it had relatively short wings, the wingspan was about twenty centimetres; its weight has been estimated at fifteen to twenty grammes. It bore a single claw on each wing. The preserved axial column length is eighty-seven millimetres. Its ribcage was not strengthened by ossified
uncinate processes but cartilaginous processes were likely present.
The ''Las Hoyas Unit 3'' site was once a forest surrounding a lake; the climate was warm with a distinct dry season. Since the skull is not known, the diet of ''Iberomesornis'' remains a subject of guesswork. It perhaps hunted insects and other small animals, plucking them out of the air or from the ground. It may have preferred to live near lakes, catching insects from the water's surface. When resting, it used its strong, clawed feet to perch on branches; the toes are long with curved claws and a low and long
hallux
Toes are the digits (fingers) of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being '' digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being '' pl ...
improved the grasping ability.
Phylogeny
''Iberomesornis romerali'' was more derived physically than ''
Archaeopteryx
''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird''), is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaīos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
''. The basal characteristic of a long tail had been lost and the
coracoid was longer. The vertebrate column of ''Iberomesornis'' ended in a
pygostyle: a splint of bone made of fused tail vertebrae that supports the tail feathers. The shorter tail and the higher position of the shoulder indicated by the strut-like coracoid — allowing for a greater wing amplitude — improved manoeuvrability, turning and swooping at speed. Flying speed may have been impaired due to the lack of a bony keel on the sternum as attachment for the flight muscles; it is uncertain whether a true ''
'' was present, through which a tendon could elevate and
supinate
Motion, the process of movement, is described using specific anatomical terms. Motion includes movement of organs, joints, limbs, and specific sections of the body. The terminology used describes this motion according to its direction relative ...
the wing.
In 1992 ''Iberomesornis'' was assigned to the
Iberomesornithidae
Iberomesornithidae is an extinct family of enantiornithine birds.
*Order Iberomesornithiformes
**Family Iberomesornithidae
***Enantiornithes gen. et sp. indet. CAGS-IG-07-CM-001
***''Iberomesornis'' (Early Cretaceous)
***''Noguerornis
''Nogue ...
. At that time it was considered to possibly have been very basal, outside of the
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Ornithothoraces
Ornithothoraces is a group of avialans that includes all enantiornithes ("opposite birds") and the euornithes ("true birds"), which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to ...
, as is reflected by its generic name. However, since the discovery of ''Iberomesornis'', many more bird fossils have come to light in the quarries of
Liaoning
Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmost ...
province, China. Well preserved specimens of Chinese species such as ''
Sinornis
''Sinornis'' is a genus of enantiornithean birds from the Lower Cretaceous Jiufotang Formation of the People's Republic of China.
When it was described in 1992, this 120 million-year-old sparrow-sized skeleton represented a new avian sharing "pr ...
'' and ''
Confuciusornis'' have permitted scientists to better understand the birds' early fossil history. Today, ''Iberomesornis'' is seen as positioned within Ornithothoraces, as a basal member of the
Enantiornithes
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
.
References
Sources
*
{{Taxonbar, from=Q134376
Enantiornitheans
Bird genera
Cretaceous birds of Europe
Cretaceous Spain
Fossils of Spain
La Huérguina Formation
Fossil taxa described in 1992
Taxa named by José Bonaparte