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The Heliomonadida (formerly Dimorphida) are a small group of
heliozoa Heliozoa, commonly known as sun-animalcules, are microbial eukaryotes ( protists) with stiff arms (axopodia) radiating from their spherical bodies, which are responsible for their common name. The axopodia are microtubule-supported projections fr ...
n
amoeboid An amoeba (; less commonly spelled ameba or amœba; plural ''am(o)ebas'' or ''am(o)ebae'' ), often called an amoeboid, is a type of cell or unicellular organism with the ability to alter its shape, primarily by extending and retracting pseudopo ...
s that are unusual in possessing
flagella A flagellum (; ) is a hairlike appendage that protrudes from certain plant and animal sperm cells, and from a wide range of microorganisms to provide motility. Many protists with flagella are termed as flagellates. A microorganism may have fro ...
throughout their life cycle.


Classification

Genetic studies place them among the
Cercozoa Cercozoa is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead defined by molecular phylogenies of rRNA and actin or polyubiquitin. They were the first major euk ...
, a group including various other
flagellate A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and thei ...
s that form filose pseudopodia. This order has recently been placed into the new
class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differently ...
of naked
filose Filose can refer to a few things: *Filose, describing members of the Filosa subphylum of Cercozoa Cercozoa is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead ...
cercozoans called Granofilosea. There are two genera in this
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
: * '' Heliomorpha'', a tiny organism found in freshwater * the larger '' Tetradimorpha'', which is distinguished by having four rather than two flagella.


Morphology

Bundles of
microtubule Microtubules are polymers of tubulin that form part of the cytoskeleton and provide structure and shape to eukaryotic cells. Microtubules can be as long as 50 micrometres, as wide as 23 to 27  nm and have an inner diameter between 1 ...
s, typically in square array, arise from a body near the flagellar bases and support the numerous axopods that project from the cell surface. Dimorphids have a single
nucleus Nucleus ( : nuclei) is a Latin word for the seed inside a fruit. It most often refers to: * Atomic nucleus, the very dense central region of an atom *Cell nucleus, a central organelle of a eukaryotic cell, containing most of the cell's DNA Nucl ...
, and
mitochondria A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used ...
with tubular cristae.


References

{{Taxonbar, from=Q5705752 Granofilosea Cercozoa orders Amoeboids Taxa named by Thomas Cavalier-Smith