Hypericum Huber-morathii
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''Hypericum huber-morathii'' is a species of flowering plant in the St John's wort family Hypericaceae. It is a small
perennial herb A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
with few stems. It has narrow and brittle stems, thick leaves, flowers in clusters of varying numbers, small yellow petals, around twenty
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s, and three styles. ''H. huber-morathii'' is closely related to '' H. minutum'' and '' H. sechmenii,'' and also shares characteristics with '' H. lanuginosum.'' The plant is endemic to Turkey, and is found among limestone rocks in a limited region of southwestern Anatolia. Originally excluded from a comprehensive monograph of '' Hypericum'', the species' placement within the genus is unclear. It has been placed in both section '' Adenosepalum'' and section '' Origanifolium''.


Etymology

The genus name ''Hypericum'' is possibly derived from the Greek words (above) and (picture), in reference to the tradition of hanging the plant over religious icons in the home. The Swiss botanist
Arthur Huber-Morath Arthur is a common male given name of Brythonic origin. Its popularity derives from it being the name of the legendary hero King Arthur. The etymology is disputed. It may derive from the Celtic ''Artos'' meaning “Bear”. Another theory, more wi ...
was the first to collect the species, and the
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''huber-morathii'' is in reference to him. In Turkish, the plant is known as .


Description

''Hypericum huber-morathii'' is a small perennial herb that grows 8–15 centimeters tall. It is entirely hairless, with a few stems that grow upright from a mostly flat base. They usually do not branch out beneath the flower clusters. The stems are narrow, roughly cylindrical, brittle in texture, and lack glands. The leaves have a very short stalk or are almost directly attached to the stem. The leaf blades are 0.5–0.9 cm long by 0.4–0.6 cm wide and are the shape of an oblong to wide oval or a more triangular oval. They are rather thick, with a rounded tip and a flat or wide wedge-shaped base. There are many pale glands on the surface of the leaf, while there are dense clusters of black glands along its edges. The flowers are in clusters that vary in number widely. There are usually 3–12 per cluster, but up to 30 have been observed. The cluster can vary in shape from somewhat corymb-like to a wide pyramid or rarely more cylindrical. The flowers are roughly 0.8 cm wide and grow from two or three nodes. The leaf-like
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s are oblong and have black glands. The sepals are around 0.25 cm long and 1.5 cm wide; they are all the same size and overlap one another. They have many point-shaped black glands on their surface and edges. The plant's petals are small and yellow, with a blunt lance shape. They have one or two black glands on their edges. Each flower has 18–20
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and 3 styles, and its
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
is 0.15 cm long. ''Hypericum huber-morathii'' can be told apart from its closest relatives, ''H. sechmenii'' and ''H. minutum,'' by several characteristics. These include its longer stems, greater number of flowers per cluster, more pointed sepals, black instead of amber glands on its petals, and earlier flowering period in June. In general, the plant has an appearance that is between that of '' H. lanuginosum'' and ''H. minutum''.''''


Taxonomy

The species was first described as ''Hypericum huber-morathii'' by Norman Robson in 1967 in the journal '' Notes from the Royal Botanic Garden, Edinburgh.'' It was originally excluded in error from a comprehensive
monograph A monograph is a specialist work of writing (in contrast to reference works) or exhibition on a single subject or an aspect of a subject, often by a single author or artist, and usually on a scholarly subject. In library cataloging, ''monograph ...
of the genus ''Hypericum'' by Robson, along with the related species '' H. formosissimum'' and '' H. minutum''. A later edition in 1996 corrected the mistake and discussed the morphology and relationships of ''H. huber-morathii''. However, it introduced ambiguity as to which section of ''Hypericum'' the species belonged. Robson acknowledged that it and the two other excluded species should have been included in his treatment of sect. ''Adenosepalum'', but then advocated for their inclusion in sect. ''Origanifolia'' based on the structure of their vittae, which would relate them most closely to '' Hypericum aviculariifolium'' in the latter section. According to Robson, removing the three species, along with several others related to '' Hypericum elodeoides'', would lead to a "purified" sect. ''Adenosepalum'' forming a "natural group" of species. Further complicating the contradiction was the species' later inclusion by Norman Robson and his colleague David Pattinson within a "''Huber-morathii'' group" inside sect. ''Adenosepalum'' in the online edition of the monograph in 2013. Under that classification, the placement of ''Hypericum huber-morathii'' was summarized as follows: '' Hypericum'' : ''Hypericum'' subg. ''Hypericum'' :: ''Hypericum'' sect. ''Adenosepalum'' ::: ''Huber-morathii'' group :::: '' H. decaisneanumH. formosissimum – H. huber-morathii'' – '' H. minutumH. sechmenii'' The name was accepted again in 2010 in a cladistic analysis of the genus, and in a review of the taxonomy of ''Hypericum'' by Robson and Sara Crockett in 2011, though no clarification was given as to its classification within the genus in either report.


Distribution, habitat, and ecology

''Hypericum huber-morathii'' is one of eight species of ''Hypericum'' sect. ''Adenosepalum'' that are native to Turkey. Its holotype was collected 19 kilometers west of
Korkuteli Korkuteli is a district of Antalya Province in the Mediterranean region of Turkey, north-west of the city of Antalya. It was previously called Istanoz or Stenez Etymology Its modern name comes from Korkut, an Ottoman prince, who was murdered by ...
in Antalya province of Anatolia''.'' The species' habitat is among limestone rocks at elevations of 1,200–1,250 meters, where it is found in association with '' Centaurea werneri'', another Turkish perennial herb. A 1997 plant registry by the World Conservation Monitoring Centre listed the species as a "single-country endemic" to Turkey that was considered Rare. Propagation of ''Hypericum huber-morathii'' is undertaken by planting seeds in the spring, barely covering them in soil. They are then allowed to germinate for 1–3 months at a temperature of 10–16 °C. The plants grow best in sunny, dry rock crevices with protection from winter dampness. Division is done in the spring, while cuttings are taken in the late summer.


References


Bibliography

* * * * * * * * {{Taxonbar, from=Q17566079 huber-morathii Flora of Turkey Plants described in 1967