''Hylocomium splendens'', commonly known as glittering woodmoss, splendid feather moss, stairstep moss, and mountain fern moss, is a

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

clonal

moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...

with a widespread distribution in

Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...

boreal forests. It is commonly found in

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

and

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

, where it is often the most abundant moss species. It also grows in the

Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...

tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...

and further south at higher elevations in, for example, northern

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

, western

Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...

East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

and the

West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...

. In Scotland it is a characteristic species of the Caledonian Forest. Under the UK's national vegetation classification system, pinewood community W18 is named as "

Pinus sylvestris ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and or ...

-Hylocomium splendens woodland", indicating its significance in this ecosystem."Hylocomium splendens: Mountain Fern Moss"
Rook.Org. Retrieved 16 May 2008.
Trees for Life. Retrieved 17 May 2008.
borealforest.org. Retrieved 17 May 2008.

Morphology

It is generally olive green, yellowish or reddish green in colour, with reddish stems and branches. These often form branches up to 20 cm long, with current year's growth starting from near the middle of the previous year's branch. This produces feathery fronds in steps. It is possible to estimate the age of a plant by counting the steps - a new level being produced each year. This form of growth enables the species to "climb" over other mosses and forest debris that falls on it. It is shade-loving, grows in soil and humus and on decaying wood and often forms mats with living parts growing on top of older, dead or dying sections. Further south, the plants are larger with several steps; further north, in the arctic

, the plants are smaller with few steps.

Ecology

Occurring widely in the

boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...

s, this plant is often found on forest floors even in relatively harsh northern latitudes. In Canada, for example, according to C. Michael Hogan the black spruce/

feathermoss Hypnales is the botanical name of an order of Bryophyta or leafy mosses. This group is sometimes called feather mosses, referring to their freely branched stems. The order includes more than 40 families and more than 4,000 species, making them ...

climax forest often occurs with moderately dense canopy featuring a forest floor of feathermosses that include ''H. splendens'', ''

Pleurozium schreberi ''Pleurozium schreberi'', the red-stemmed feathermoss or Schreber's big red stem moss, is a moss with a loose growth pattern. The root name ''pleuro'' comes from the Latin for ribs, possibly describing how the parts branch from the stem. The spe ...

'' and ''

Ptilium crista-castrensis ''Ptilium crista-castrensis'', the knights plume moss or ostrich-plume feathermoss, is a moss species within the family Pylaisiaceae, in the class Bryopsida, subclass Bryidae and order Hypnales. Ecology This species occurs on the floor of Cana ...

''. Experiments simulating global change with increased nutrient deposits and warming indicate that increased nutrient deposits and climate warming may have negative impact on growth of Hylocomium splendens.

Use

The species has a commercial use in floral exhibitions and for lining fruit and vegetable storage boxes. In the past it was utilised as a floor covering for dirt floors. In Alaska and northern Canada it is still used for filling the gaps between the logs in log cabins. It has anti-bacterial qualities and may also contain anti-

tumour A neoplasm () is a type of abnormal and excessive growth of tissue. The process that occurs to form or produce a neoplasm is called neoplasia. The growth of a neoplasm is uncoordinated with that of the normal surrounding tissue, and persists ...

agents.

References

External links

''Hylocomium splendens'' Images
{{Taxonbar, from=Q149101 Hypnales Flora of North America Flora of Northern Europe