The Shasta salamander (''Hydromantes shastae'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten ...

in the family

Plethodontidae Plethodontidae, or lungless salamanders, are a family of salamanders. Most species are native to the Western Hemisphere, from British Columbia to Brazil, although a few species are found in Sardinia, Europe south of the Alps, and South Korea. In ...

. It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Shasta County Shasta County (), officially the County of Shasta, is a county in the northern portion of the U.S. state of California. Its population is 182,155 as of the 2020 census, up from 177,223 from the 2010 census. The county seat is Redding. Shasta ...

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

Taxonomy

Formerly considered to be a single species with a rather small distribution in the vicinity of

Shasta Lake Shasta Lake, also popularly known as Lake Shasta, is a reservoir in Shasta County, California, United States. It began to store water in 1944 due to the impounding of the Sacramento River by Shasta Dam, the ninth tallest dam in the United State ...

, a 2018 study found it to comprise three species, including ''H. shastae sensu stricto'', and two new species: the Samwel Shasta salamander (''H. samweli''), found in the north-central and northwest sides of the lake, and the Wintu Shasta salamander (''H. wintu'') on the northern shore of the lake near the

McCloud River The McCloud River is a longU.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed March 10, 2011 river that flows east of and parallel to the upper Sacramento River, in Siskiyou County and Sh ...

. Both species are physically similar to ''H. shastae'' and can only be distinguished by range and genetics.

Distribution and habitat

It is found in the

Cascade Range The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as the North Cascades, ...

in areas around

with

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

substrate, primarily between Squaw Creek and the

Pit River The Pit River is a major river draining from northeastern California into the state's Central Valley. The Pit, the Klamath and the Columbia are the only three rivers in the U.S. that cross the Cascade Range. The longest tributary of the Sacr ...

, but also on the south side of Shasta Lake and in the vicinity of

Ingot An ingot is a piece of relatively pure material, usually metal, that is cast into a shape suitable for further processing. In steelmaking, it is the first step among semi-finished casting products. Ingots usually require a second procedure of sh ...

. Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are temperate

forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...

s, freshwater

spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a ...

s, rocky areas, and

cave A cave or cavern is a natural void in the ground, specifically a space large enough for a human to enter. Caves often form by the weathering of rock and often extend deep underground. The word ''cave'' can refer to smaller openings such as sea ...

Threats

Its small distribution was likely fragmented by the creation of the lake and the construction of the

Shasta Dam Shasta Dam (called Kennett Dam before its construction) is a concrete arch-gravity dam across the Sacramento River in Northern California in the United States. At high, it is the eighth-tallest dam in the United States. Located at the north e ...

. Proposals to raise the water levels of the lake would likely flood more habitats and further threaten it. In addition, the splitting of this species into three distinct species with highly restricted ranges likely makes it even more endangered previously thought. However, in 2021 the

U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with oth ...

found the species to not warrant

Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...

protections.

References

Amphibians of the United States Hydromantes Endemic fauna of California Amphibians described in 1953 Taxonomy articles created by Polbot {{Plethodontidae-stub