Hydrocotyle Acutiloba
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''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-
aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
s formerly classified in the family Apiaceae, now in the family Araliaceae.


Description

Water pennyworts, ''Hydrocotyles'', are very common. They have long creeping stems that often form dense mats, often in and near ponds, lakes, rivers, and marshes, and some species in coastal areas by the sea. ;Leaves: Simple, with small leafy outgrowth at the base, kidney shaped to round. Leaf edges are scalloped. The leaf surfaces of ''Hydrocotyle'' are prime grounds for
oviposition The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
of many butterfly species, such as '' Anartia fatima''. ;Flowers: Flower clusters are simple and flat-topped or rounded.
Involucral bracts In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, ...
at the base of each flower. Indistinct
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
. ;Fruits and reproduction: Elliptical to round with thin ridges and no oil tubes (vitta) which is characteristic in the fruit of umbelliferous plants. :The prostrate plants reproduce by seed and by sending roots from stem nodes.


Selected species

The genus ''Hydrocotyle'' has between 75 and 100 species that grow in tropical and temperate regions worldwide. A few species have entered the world of cultivated ornamental aquatics. A list of selected species: *'' Hydrocotyle americana'' L. — American marshpennywort *''Hydrocotyle asiatica'' L. is now correctly known as '' Centella asiatica'' *'' Hydrocotyle batrachium'' Hance *'' Hydrocotyle benguetensis'' Elm. *'' Hydrocotyle bonariensis'' Lam. — largeleaf pennywort *'' Hydrocotyle bowlesioides'' Mathias & Constance — largeleaf marshpennywort *'' Hydrocotyle calcicola'' *'' Hydrocotyle dichondroides'' Makino *'' Hydrocotyle dielsiana'' *'' Hydrocotyle heteromeria'' — waxweed *''
Hydrocotyle hexagona ''Hydrocotyle hexagona'' is a species of flowering plant in the family Araliaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forests. It is threatened by habitat loss Habitat destruction (also te ...
'' *'' Hydrocotyle himalaica'' *'' Hydrocotyle hirsuta'' Sw. — yerba de clavo *''
Hydrocotyle hirta ''Hydrocotyle hirta'', commonly known as the hairy pennywort, is a species of flowering plant in the family Araliaceae native to Australia. References hirta Hirta ( gd, Hiort) is the largest island in the St Kilda archipelago, on the wes ...
'' *''
Hydrocotyle hitchcockii ''Hydrocotyle hitchcockii'' is a species of flowering plant in the family Araliaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist lowland forests and subtropical or tropical moist montane forests. It is threat ...
'' *'' Hydrocotyle hookeri'' *''Hydrocotyle hydrophila'' - endemic to New Zealand *''
Hydrocotyle javanica ''Hydrocotyle javanica'', commonly known as Java pennywort, is a species of ''Hydrocotyle''. It is a prostrate herb found in NE India and SE Asia. Leaves are simple, circular-heart-shaped, with seven triangular shallow lobes. Leaves are 2.5-5 x 3 ...
'' Thunb. *''
Hydrocotyle keelungensis ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' Liu, Chao & Chuang *''
Hydrocotyle leucocephala ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the ...
'' Cham. & Schltdl. — Brazilian pennywort *''
Hydrocotyle mannii ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the ...
'' Hook.f. *''
Hydrocotyle microphylla ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' A.Cunn. *''
Hydrocotyle moschata ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' G. Forst. — musky marshpennywort *'' Hydrocotyle nepalensis'' Hook. *''
Hydrocotyle novae-zelandiae ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' DC. *''
Hydrocotyle phoenix ''Hydrocotyle phoenix'' is a species of annual pennywort and is commonly called fire pennywort. It is only known to grow in south-west Australia, specifically in fire prone habitats; it is unique in this regard as it is the only species of ''Hyd ...
'' A.J. Perkins — fire pennywort *''
Hydrocotyle prolifera ''Hydrocotyle prolifera'', commonly called whorled marshpennywort , is a species of flowering plant in the ginseng family (Araliaceae The Araliaceae are a family of flowering plants composed of about 43 genera and around 1500 species consisti ...
'' Kellogg — whorled marshpennywort *''
Hydrocotyle pseudoconferta ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' *''
Hydrocotyle pusilla ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' A. Rich. — tropical marshpennywort *''
Hydrocotyle ramiflora ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the ...
'' *''
Hydrocotyle ranunculoides ''Hydrocotyle ranunculoides'', known commonly as floating pennywort, or floating marshpennywort, is an aquatic plant in the family Apiaceae. It is native to North and South America. Origin and invasiveness Water pennywort is an aquatic plant, ...
'' L. f. — floating marshpennywort, floating marshpennywort, floating pennyroyal *''
Hydrocotyle salwinica ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' *''
Hydrocotyle setulosa ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the f ...
'' Hayata *''
Hydrocotyle sibthorpioides ''Hydrocotyle sibthorpioides'' is a small plant native to southeastern Asia. It is also referred to as lawn marshpennywort. It is a dicot, traditionally placed in the family Apiaceae, but more recently suggested to belong in the Araliaceae. I ...
'' Lam. — lawn marshpennywort *''
Hydrocotyle tambalomaensis ''Hydrocotyle tambalomaensis'' is a species of flowering plant in the family Araliaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist montane forests and subtropical or tropical high-altitude grassland. It is t ...
'' *''
Hydrocotyle tripartita ''Hydrocotyle tripartita'' is a perennial herb flowering with small white flowers that comes from New Zealand and the Australian states of Queensland, New South Wales and Victoria Victoria most commonly refers to: * Victoria (Australia), a st ...
'' *''
Hydrocotyle umbellata ''Hydrocotyle umbellata'' is an aquatic plant that thrives in wet, sandy habitat. Its English common name is manyflower marshpennywort or dollarweed. It is native to North America and parts of South America. In Brazil it is known as acariçoba a ...
'' L. — manyflower marshpennywort, umbrella pennyroyal *''
Hydrocotyle verticillata ''Hydrocotyle verticillata'', also known as whorled pennywort, whorled marshpennywort or shield pennywort, is a flowering plant found in South and North America and the West Indies. Its creeping habit and unusual leaves give it its common names. ...
'' Thunb. — whorled marshpennywort, whorled marshpennywort, whorled pennyroyal *'' Hydrocotyle vulgaris'' L. — marsh pennywort, common pennywort *''
Hydrocotyle wilfordii ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the ...
'' *''
Hydrocotyle wilsonii ''Hydrocotyle'', also called floating pennywort, water pennywort, Indian pennywort, dollar weed, marsh penny, thick-leaved pennywort and even white rot is a genus of prostrate, perennial aquatic or semi-aquatic plants formerly classified in the ...
'' *''
Hydrocotyle yanghuangensis ''Hydrocotyle yanghuangensis'' is a species of flowering plant in the family Araliaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist montane forests and subtropical or tropical high-altitude grassland. It is t ...
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Distribution

''Hydrocotyleae'' grow in wet and damp places in the tropics and the temperate zones.


Fossil record

One fossil fruit of a ''Hydrocotyle'' sp. has been extracted from borehole samples of the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million y ...
fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) zczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska) Acta Palaeobotanica 1979 20 (1): 3-117.


References


External links

* * {{Authority control Apiales genera Taxa named by Carl Linnaeus Aquatic plants