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A hydathode is a type of pore, commonly found in
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s, that secretes water through pores in the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rele ...
or leaf margin, typically at the tip of a marginal tooth or serration. Hydathodes occur in the leaves of submerged aquatic plants such as ''
Ranunculus fluitans ''Ranunculus fluitans'' (the river water-crowfoot,) is a species of buttercup. It is a perennial water plant, which when in favourable conditions (such as fast flowing water,) can grow up to 6 m (20 ft) height.J. S. Rodwell; ''British P ...
'' as well as herbaceous plants of drier habitats such as '' Campanula rotundifolia''. They are connected to the plant vascular system by a vascular bundle. Hydathodes are commonly seen in
water lettuce ''Pistia'' is a genus of aquatic plants in the arum family, Araceae. It is the sole genus in the tribe ''Pistieae'' which reflects its systematic isolation within the family. The single species it comprises, ''Pistia stratiotes'', is often cal ...
,
water hyacinth ''Pontederia crassipes'' (formerly ''Eichhornia crassipes''), commonly known as common water hyacinth is an aquatic plant native to South America, naturalized throughout the world, and often invasive outside its native range.rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
, balsam, and many other species. Hydathodes are made of a group of living cells with numerous intercellular spaces filled with water, but few or no chloroplasts, and represent modified bundle-ends. These cells (called ''epithem cells'') open out into one or more sub-epidermal chambers. These, in turn, communicate with the exterior through an open water stoma or open pore. The water stoma structurally resembles an ordinary
stoma In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
, but is usually larger and has lost the power of movement. Hydathodes are involved in the process of guttation, in which positive xylem pressure (due to
root pressure Root pressure is the transverse osmotic pressure within the cells of a root system that causes sap to rise through a plant stem to the leaves. Root pressure occurs in the xylem of some vascular plants when the soil moisture level is high either ...
) causes liquid to exude from the pores. Some halophytes possess
gland In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
ular
trichome Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a pla ...
s that actively secrete salt in order to reduce the concentration of cytotoxic inorganic ions in their cytoplasm; this may lead to the formation of a white powdery substance on the surface of the leaf. Hydathodes are of two types: # passive hydathodes, formed when a leaf vein terminates in an epithem (an area of thin-walled
parenchyma Parenchyma () is the bulk of functional substance in an animal organ or structure such as a tumour. In zoology it is the name for the tissue that fills the interior of flatworms. Etymology The term ''parenchyma'' is New Latin from the word π ...
). # active hydathodes, formed when epidermal cells lose water actively.


See also

*
Transpiration Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Water is necessary for plants but only a small amount of water taken up by the roots is used for growth a ...
*
Vascular plants Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
* Xylem


References


External links


hydathode physiology
Plant anatomy {{botany-stub