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''Humbertium'' is a genus of
land planarian Geoplanidae is a family of flatworms known commonly as land planarians or land flatworms. These flatworms are mainly predators of other invertebrates, which they hunt, attack and capture using physical force and the adhesive and digestive proper ...
s of the subfamily
Bipaliinae Bipaliinae is a subfamily of land planarians found mainly in Madagascar, the Indian subcontinent and Southeast Asia, although some species have been introduced worldwide. Description The subfamily Bipaliinae is characterized by having a semi ...
(hammerhead flatworms).


Description

Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to the one in ''
Bipalium ''Bipalium'' is a genus of large predatory land planarians. They are often loosely called "hammerhead worms" or "broadhead planarians" because of the distinctive shape of their head region. Land planarians are unique in that they possess a "creep ...
'', but with the ovovitelloducts entering the female atrium anteriorly and not posteriorly as in ''Bipalium''.


Etymology

The genus ''Humbertium'' is named after the Swiss naturalist
Aloïs Humbert Aloïs Humbert (22 September 1829, in Geneva – 14 May 1887) was a Swiss naturalist and paleontologist who specialized in the study of myriapods. He also described new vertebrates (fishes, reptiles, mammals), molluscs and flatworms. In 1852 he ...
who described several species from
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
that are now classified in the genus.


Species

The genus ''Humbertium'' contains the following species: *''
Humbertium core ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (de Beauchamp, 1930) *''
Humbertium covidum ''Humbertium covidum'' is a species of predatory land flatworm, found in France and Italy. The Holotype specimen is MNHN JL351B. Description ''H. covidum'' is a small hammerhead flatworm, about in length. The dorsal face is completely metalli ...
'' Justine, Gastineau, Gros, Gey, Ruzzier, Charles & Winsor, 2022 *''
Humbertium depressum ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Ritter-Záhony, 1905) *''
Humbertium diana ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Humbert, 1862) *''
Humbertium dodabettae ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (de Beauchamp, 1930) *''
Humbertium ferruginoideum ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Sabussowa, 1925) *''
Humbertium kelleri ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (von Graff, 1899) *''
Humbertium longicanale ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Sabussowa, 1925) *''
Humbertium palnisium ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (de Beauchamp, 1930) *''
Humbertium penangense ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Kawakatsu, 1986) *''
Humbertium penrissenense ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (de Beauchamp, 1925) *''
Humbertium phebe ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Humbert, 1862) *''
Humbertium proserpina ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (Humbert, 1862) *''
Humbertium pseudophallicum ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (de Beauchamp, 1925) *''
Humbertium ravenalae ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the copulatory apparatus, similar to ...
'' (von Graff, 1899) *''
Humbertium subboreale ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the reproductive system of planarians ...
'' (Sabussowa, 1925) *''
Humbertium umbrinum ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the reproductive system of planarians ...
'' (Geba, 1909) *''
Humbertium voigti ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the reproductive system of planarians ...
'' (von Graff, 1899) *''
Humbertium woodworthi ''Humbertium'' is a genus of land planarians of the subfamily Bipaliinae (hammerhead flatworms). Description Species of ''Humbertium'' are characterized by the presence of a well developed penis papilla in the reproductive system of planarians ...
'' (von Graff, 1899)


References

Geoplanidae Rhabditophora genera {{flatworm-stub