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The many-horned adder (''Bitis cornuta'') is a
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. It is found in certain rocky
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
areas, mostly along the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
coast of southern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, in western South Africa and southwestern Namibia. It has characteristic tufts of "horns" above each eye.Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . Like all other vipers, it is
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
. No
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are currently recognized.


Taxonomy

Common names include many-horned adder, hornsman, western hornsman adder, and western many-horned adder.Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . The type locality given is ''"Cap de Bonne-espérance"'' (Cape of Good Hope, South Africa). Actually, according to Patterson's itinerary, the type was observed in coastal
Namaqualand Namaqualand (khoekhoe: "Nama-kwa" meaning Nama Khoe people's land) is an arid region of Namibia and South Africa, extending along the west coast over and covering a total area of . It is divided by the lower course of the Orange River into ...
, on 1 September 1779.


Description

Small and stout, it grows to a typical total length (body and tail) of 30–50 cm (about 12–20 inches). The maximum recorded total length is for a captive specimen. Two to five raised, horn-like scales occur above each eye. Boulenger GA. 1896. ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., Containing the...Viperidæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers.) xiv + 727 pp. + Plates I.- XXV. (''Bitis cornuta'', pp. 497-498.)
Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, it has a grey to reddish-brown ground colour, overlaid with four longitudinal series of large, dark-brown blotches, which are squarish or parallelogram-like in shape, and edged with white.
Ventral Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Th ...
ly, it is whitish or tan, either uniform or speckled with dark brown. On the dorsal surface of the head, dark, symmetrical markings may form an arrowhead.


Distribution and habitat

The many-horned adder ranges from the coastal region of southwest
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
through west and southwest
Cape Province The Province of the Cape of Good Hope ( af, Provinsie Kaap die Goeie Hoop), commonly referred to as the Cape Province ( af, Kaapprovinsie) and colloquially as The Cape ( af, Die Kaap), was a province in the Union of South Africa and subsequen ...
in
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, with a few isolated populations in eastern Cape Province. This species prefers rocky desert areas in dwarf
succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
veld Veld ( or ), also spelled veldt, is a type of wide open rural landscape in :Southern Africa. Particularly, it is a flat area covered in grass or low scrub, especially in the countries of South Africa, Lesotho, Eswatini, Zimbabwe and Bots ...
and
mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited Summit (topography), summit area, and ...
slopes in
heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a cooler ...
vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
.


Behavior

With a nervous disposition, when disturbed, it will hiss loudly and strike so energetically that most of its body is lifted off the ground in the process. However, it usually settles down in captivity.


References


Further reading

* Branch, Bill. 2004. ''Field Guide to Snakes and Other Reptiles of Southern Africa''. Sanibel Island, Florida: Ralph Curtis Books. 399 pp. . (''Bitis cornuta'', pp. 116–117 + Plate 13.) * Daudin FM. 1803. ''Histoire Naturelle, Générale et Particulière des Reptiles; Ouvrage faisant suite à l'Histoire Naturelle générale et particulière, composée par LECLERC DE BUFFON; et rédigée par C.S. SONNINI, membre de plusieurs Sociétés savantes. Tome VI.'' Paris: F. Dufart. 447 pp. (''Vipera cornuta'', p. 188.)


External links


Many-horned adder (''Bitis cornuta'')
a
ARKive
Accessed 5 October 2006. * . Accessed 1 March 2007. {{Taxonbar, from=Q2661334 Bitis Snakes of Africa Reptiles of Namibia Reptiles of South Africa Reptiles described in 1803 Taxa named by François Marie Daudin