Hormogonia

Hormogonia are motile filaments of cells formed by some

cyanobacteria Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, bl ...

in the order Nostocales and Stigonematales. They are formed during vegetative reproduction in unicellular, filamentous

, and some may contain heterocysts and akinetes. Cyanobacteria differentiate into hormogonia when exposed to an

environmental stress Stress, either physiological, biological or psychological, is an organism's response to a stressor such as an environmental condition. Stress is the body's method of reacting to a condition such as a threat, challenge or physical and psyc ...

or when placed in new media. Hormogonium differentiation is crucial for the development of nitrogen-fixing plant cyanobacteria symbioses, in particular that between cyanobacteria of the genus ''

Nostoc ''Nostoc'', also known as star jelly, troll’s butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch’s jelly, is the most common genus of cyanobacteria found in var ...

'' and their hosts. In response to a hormogonium-inducing factor (HIF) secreted by plant hosts, cyanobacterial symbionts differentiate into hormogonia and then dedifferentiate back into vegetative cells after about 96 hours. Hopefully, they have managed to reach the plant host by this time. The bacteria then differentiate specialized nitrogen-fixing cells called heterocysts and enter into a working symbiosis with the plant. Depending on species, Hormogonia can be many hundreds of micrometers in length and can travel as fast as 11 μm/s. They move via

gliding motility Gliding motility is a type of translocation used by microorganisms that is independent of propulsive structures such as flagella, pili, and fimbriae. Gliding allows microorganisms to travel along the surface of low aqueous films. The mechanisms of ...

, requiring a wettable surface or a viscous substrate, such as

agar Agar ( or ), or agar-agar, is a jelly-like substance consisting of polysaccharides obtained from the cell walls of some species of red algae, primarily from ogonori (''Gracilaria'') and "tengusa" (''Gelidiaceae''). As found in nature, agar i ...

for motion. Cell anatomy Cyanobacteria {{cyanobacteria-stub