Hoploclonia Gecko
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''Hoploclonia gecko'' is a relatively small, spiny and darkly colored
stick insect The Phasmatodea (also known as Phasmida, Phasmatoptera or Spectra) are an order of insects whose members are variously known as stick insects, stick-bugs, walking sticks, stick animals, or bug sticks. They are also occasionally referred to as ...
species that is native to the northwest of
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
.


Taxonomy

Alfred Russel Wallace Alfred Russel Wallace (8 January 1823 – 7 November 1913) was a British naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution through natural se ...
collected in
Sarawak Sarawak (; ) is a States and federal territories of Malaysia, state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the M ...
in 1858 a number ofspecimens, but did not leave any more precise information about the location.
John Obadiah Westwood John Obadiah Westwood (22 December 1805 – 2 January 1893) was an English entomologist and archaeologist also noted for his artistic talents. He published several illustrated works on insects and antiquities. He was among the first entomologist ...
described these the following year as ''Acanthoderus gecko''. When describing it, both males and females were available to him. Because of the
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
chosen by Westwood, it is also called "Gecko Stick Insect". In 1875
Carl Stål Carl Stål (21 March 1833 – 13 June 1878) was a Swedish entomologist specialising in Hemiptera. He was born at Karlberg Castle, Stockholm on 21 March 1833 and died at Frösundavik near Stockholm on 13 June 1878. He was the son of architect, a ...
established the genus ''
Hoploclonia ''Hoploclonia'' is the only genus of the tribe Hoplocloniini and brings together relatively small and darkly coloured Phasmatodea species. Characteristics The representatives of this genus are very small with 35 to 40 mm in the male a ...
'' for this species alone, which became the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
as ''Hoploclonia gecko''. The genus remained monotypical until the description of ''
Hoploclonia cuspidata ''Hoploclonia cuspidata'' is a stick insect species native to the north of Borneo and is also called Brunei Hoploclonia stick insect. Taxonomy When Josef Redtenbacher described the species in 1906, both males and females were available to him ...
'' in 1906. In 1995 a female of
Philip Edward Bragg Philip, also Phillip, is a male given name, derived from the Greek (''Philippos'', lit. "horse-loving" or "fond of horses"), from a compound of (''philos'', "dear", "loved", "loving") and (''hippos'', "horse"). Prominent Philips who popularize ...
from the specimens collected by Wallace was selected as
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
. It is deposited together with two male paralectotypes in the
Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history collections that include current and historical records of animals, plants, fungi, ecosystems, geology, paleontology, climatology, and more. ...
. Other specimens collected by Wallace are kept as paralectotypes in the
Oxford University Museum of Natural History The Oxford University Museum of Natural History, sometimes known simply as the Oxford University Museum or OUMNH, is a museum displaying many of the University of Oxford's natural history specimens, located on Parks Road in Oxford, England. It a ...
.


Description

The insects, wingless in both sexes, have typical spines on their bodies like all representatives of this genus. In the long males, these are located in pairs on the head and the
thorax The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the cre ...
, but not on the
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
. The pair of spines on the
pronotum The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum (dorsal), the prosternum (ventral), and the propleuron (lateral) on ea ...
is forked.
Adult An adult is a human or other animal that has reached full growth. In human context, the term ''adult'' has meanings associated with social and legal concepts. In contrast to a " minor", a legal adult is a person who has attained the age of major ...
males are dominated by a dark, mostly black-brown basic color with a bright yellow-orange to red longitudinal line, which extends from the rear edge of the pronotum to the middle of the abdomen. As with the other representatives of the genus, the areas around the coxae, as well as the
distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
ends of the
femurs The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with t ...
and the
proximal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
ends of the tibae, ie the knee area, colored light yellow to orange. The females are long. They can be colored more or less high-contrast light, dark or red-brown. They have clear spines on the thorax. On the
mesothorax The mesothorax is the middle of the three segments of the thorax of hexapods, and bears the second pair of legs. Its principal sclerites (exoskeletal plates) are the mesonotum (dorsal), the mesosternum (ventral), and the mesopleuron (lateral) on ...
the first pair of spines forms a flat triangle, very broad at the base, which is much more pronounced than in other ''Hoploclonia'' species. Overall, they are very robust from the habitus. The relatively short abdomen is plump in egg-laying females and approximately cylindrical in cross-section. The end of the abdomen is formed by a short
ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
, which is used to lay the eggs in the ground.


Distribution

''Hoploclonia gecko'' is native to the northwest of Borneo, more precisely in the far west of the
Malaysian Malaysian may refer to: * Something from or related to Malaysia, a country in Southeast Asia * Malaysian Malay, a dialect of Malay language spoken mainly in Malaysia * Malaysian people, people who are identified with the country of Malaysia regard ...
state
Sarawak Sarawak (; ) is a States and federal territories of Malaysia, state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the M ...
. The species was found here in
Bako National Park Bako National Park is a national park in Kuching Division, Sarawak, Malaysia. Established in 1957, it is the oldest national park in Sarawak. It covers an area of at the tip of the Muara Tebas peninsula at the mouth of the Bako and Kuching River ...
, on
Mount Santubong Mount Santubong ( ms, Gunung Santubong) is a mountain in the Malaysian state of Sarawak. It is located about 35 km north of the state capital Kuching. Biological importance In 1855 a British naturalist, Alfred Russel Wallacewho was stayi ...
, on
Mount Serapi Mount is often used as part of the name of specific mountains, e.g. Mount Everest. Mount or Mounts may also refer to: Places * Mount, Cornwall, a village in Warleggan parish, England * Mount, Perranzabuloe, a hamlet in Perranzabuloe parish, C ...
, in
Matang Matang (Jawi alphabet, Jawi: ماتڠ; ) is a mukim in Larut, Matang and Selama District, Perak, Malaysia. It has many small towns and villages and is located near Taiping, Perak, Taiping, Simpang, Perak, Simpang, Kamunting and Kuala Sepetang. ...
and Lingga. The main area of distribution is the area around
Kuching Kuching (), officially the City of Kuching, is the capital and the most populous city in the States and federal territories of Malaysia, state of Sarawak in Malaysia. It is also the capital of Kuching Division. The city is on the Sarawak River ...
, the capital of Sarawak. On Mount Santubong she was found up to a height of .


Way of life and reproduction

Like almost all stick insects, ''Hoploclonia gecko'' is nocturnal. To eat, they do not climb higher than up the food plants. The females lay their eggs with the ovipositor only a few millimeters deep in the ground. The eggs are long, high and wide. Like all ''Hoploclonia'' species, they are more arched on the dorsal side and have a lid (operculum) sloping towards the ventral side, which forms an opercular angle of less than 10 degrees. Of the three arms of the micropylar plate, one faces the lid, while the other two run laterally in the direction of the lower pole and enclose about 2/3 of the egg.


In terraristics

''Hoploclonia gecko'' was the first species of the genus that appeared in the
terrarium A terrarium (plural: terraria or terrariums) is usually a sealable glass container containing soil and plants that can be opened for maintenance to access the plants inside; however, terraria can also be open to the atmosphere. Terraria are ofte ...
at the end of the 1980s. Philip Bragg collected some specimens in December 1987 in the Bako National Park, which he was able to successfully reproduce and spread. He brought further representatives of this species with him from Mount Serapi in 1990. The species was given PSG number 110 by the Phasmid Study Group. As with all ''Hoploclonia'' species, keeping and breeding is considered difficult. Small terrariums with higher humidity and a substrate for laying eggs are required. Leaves of
bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...
, other
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...
, as well as
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
,
ivy ''Hedera'', commonly called ivy (plural ivies), is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to western, central and southern Europe, Macaronesia, northwestern Africa and ...
, ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' and ''
Pyracantha ''Pyracantha'' (from Greek "fire" and "thorn", hence firethorn) is a genus of large, thorny evergreen shrubs in the family Rosaceae, with common names firethorn or pyracantha. They are native to an area extending from Southwest Europe east to ...
'' species are accepted as food.


References

Westwood, J. O. (1859) '' Catalog of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. '' London 1859, p. 52 & Plate XXVI: 6-7
online version
Bragg, P. E. (2001). ''Phasmids of Borneo'', Natural History Publications (Borneo) Sdn. Bhd., Kota Kinabalu, Sabah, Malaysia, pp. 110-111, pp. 120-123, '' Hoploclonia gecko '' on phasmatodea.com
by Hennemann, F. H.; Conle, O. V.; Kneubühler, B. & Valero, P.
Brock, P. D.; Büscher, T. H. & Baker, E. W.
Phasmida Species File Online
'. Version 5.0. (accessdate 16 August 2021)
Phasmid Study Group Culture List
/ref>


External links

* * {{Taxonbar, from=Q1339372 Phasmatodea Phasmatodea of Asia Insects described in 1859