''Hoplestigma'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Hoplestigmataceae.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. 2007. ''Flowering Plant Families of the World''. Firefly Books: Ontario, Canada. . One

study Study or studies may refer to: General * Education **Higher education * Clinical trial * Experiment * Observational study * Research * Study skills, abilities and approaches applied to learning Other * Study (art), a drawing or series of drawi ...

pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...

suggested that ''Hoplestigma'' might be

related ''Related'' is an American comedy-drama television series that aired on The WB from October 5, 2005, to March 20, 2006. It revolves around the lives of four close-knit sisters of Italian descent, raised in Brooklyn and living in Manhattan. The ...

to the family Ehretiaceae.Joan W. Nowicke and James S. Miller. 1989. "Pollen morphology and the relationships of Hoplestigmataceae". ''Taxon'' 38(1):12-16. In a

comparison Comparison or comparing is the act of evaluating two or more things by determining the relevant, comparable characteristics of each thing, and then determining which characteristics of each are similar to the other, which are different, and t ...

chloroplast A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in ...

DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...

s in 2014, ''Hoplestigma'' formed a strongly supported

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

with ''

Coldenia ''Coldenia'', named after C. Colden, is a monotypic genus of flowering plants traditionally included in the borage family, Boraginaceae ''sensu lato''. It was assigned to the subfamily Ehretioideae, but molecular data revealed it to be more clos ...

'' and genera that have always been placed in Cordiaceae.Maximilian Weigend, Federico Luebert, Marc Gottschling, Thomas L.P. Couvreur, Hartmut H. Hilger and James S. Miller. 2014. "From capsules to nutlets — phylogenetic relationships in the Boraginales". ''Cladistics'' 30(5):508-518. . The authors of that study recommended that ''Hoplestigma'' and ''Coldenia'' be included in Cordiaceae. However, it was traditionally included in

Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single family of the order ...

'' sensu lato'', as did the

APG IV The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publishe ...

classificationPeter F. Stevens (2001 onwards). Angiosperm Phylogeny Website. At: Missouri Botanical Garden Website. (see ''External links'' below) but subsequently placed in its own family within the Boraginales

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

, by the Boraginales Working Group. ''Hoplestigma'' has two

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

. Both are rare trees of lowland

jungle A jungle is land covered with dense forest and tangled vegetation, usually in tropical climates. Application of the term has varied greatly during the past recent century. Etymology The word ''jungle'' originates from the Sanskrit word ''jaṅ ...

West West or Occident is one of the four cardinal directions or points of the compass. It is the opposite direction from east and is the direction in which the Sunset, Sun sets on the Earth. Etymology The word "west" is a Germanic languages, German ...

and

Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

.David J. Mabberley. 2008. ''Mabberley's Plant-Book''. Cambridge University Press: UK. The genus name ''Hoplestigma'' is

derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...

from the Greek , "a hoof or a cloven hoof" and ''stigma'', "a flower stigma". The

botanical name A botanical name is a formal scientific name conforming to the '' International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the ''Intern ...

is a reference to the deeply

bifid Bifid refers to something that is split or cleft into two parts. It may refer to: * Bifid, a variation in the P wave, R wave, or T wave in an echocardiogram in which a wave which usually has a single peak instead has two separate peaks * Bifid ci ...

style.Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'' volume II. CRC Press: Boca Raton; New York; Washington,DC;, USA. London, UK. (vol. II). (see ''External links'' below).

References

Bibliography

External links

CordiaceaeMissouri Botanical Garden Website

CRC World Dictionary of Plant Names: D-LBotany & Plant ScienceLife ScienceCRC Press
{{Taxonbar, from=Q135170 Boraginaceae Taxonomy articles created by Polbot