''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied, limbless invertebrates that dwell on the ocean floor and are usually

detritivore Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...

. They resemble a

cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated Vine#Horticultural climbing plants, creeping vine plant in the Cucurbitaceae family that bears usually cylindrical Fruit, fruits, which are used as culinary vegetables.

in form. The genus contains some species that are harvested and sold as food.

Species

The following species are recognised in the genus ''Holothuria'': * Subgenus ''Acanthotrapeza'' ** ''

Holothuria coluber ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria kubaryi ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria pyxis'' ** '' Holothuria tripilata'' * Subgenus ''Cystipus'' ** '' Holothuria casoae'' ** '' Holothuria cubana'' ** '' Holothuria dura'' ** '' Holothuria hartmeyeri'' ** '' Holothuria inhabilis'' ** '' Holothuria jousseaumei'' ** '' Holothuria mammosa'' ** '' Holothuria occidentalis'' ** ''

Holothuria pseudofossor ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria rigida ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria sucosa ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria turrisimperfecta ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' * Subgenus ''Halodeima'' ** ''

Holothuria atra ''Holothuria atra'', commonly known as the black sea cucumber or lollyfish, is a species of marine invertebrate in the family Holothuriidae. It was placed in the subgenus ''Halodeima'' by Pearson in 1914, making its full scientific name ''Holothu ...

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Holothuria edulis ''Holothuria edulis'', commonly known as the edible sea cucumber or the pink and black sea cucumber, is a species of echinoderm in the family Holothuriidae. It was placed in the subgenus ''Halodeima'' by Pearson in 1914, making its full scientifi ...

'' ** '' Holothuria enalia'' ** ''

Holothuria floridana ''Holothuria floridana'', the Florida sea cucumber, is a species of marine invertebrate in the Family (biology), family Holothuriidae. It is found on the seabed just below the low tide mark in Florida, the Gulf of Mexico, the Bahamas and the Cari ...

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Holothuria grisea ''Holothuria grisea'', the gray sea cucumber, is a mid-sized coastal species of sea cucumber found in shallow tropical waters of the Atlantic Ocean from Florida to Southern Brazil and West Africa. They have a variety in color and can range from r ...

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Holothuria inornata ''Holothuria'' is the type genus of the marine animal Family (biology), family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. Th ...

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Holothuria kefersteinii ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria manningi'' ** '' Holothuria mexicana'' ** ''

Holothuria nigralutea ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria pulla'' ** ''

Holothuria signata ''Holothuria'' is the type genus of the marine animal Family (biology), family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. Th ...

'' * Subgenus ''Holothuria'' ** '' Holothuria caparti'' ** '' Holothuria dakarensis'' ** '' Holothuria fungosa'' ** ''

Holothuria helleri ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria mammata ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria massaspicula'' ** '' Holothuria stellati'' ** ''

Holothuria tubulosa ''Holothuria tubulosa'', the cotton-spinner or tubular sea cucumber, is a species of sea cucumber in the family Holothuriidae. It is the type species of the genus '' Holothuria'' and is placed in the subgenus ''Holothuria'', making its full nam ...

'' * Subgenus ''Lessonothuria'' ** '' Holothuria cavans'' ** '' Holothuria cumulus'' ** ''

Holothuria duoturricula ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria glandifera ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria immobilis'' ** ''

Holothuria insignis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria lineata ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria multipilula'' ** ''

Holothuria pardalis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria tuberculata'' ** '' Holothuria verrucosa'' * Subgenus ''Mertensiothuria'' ** '' Holothuria albofusca'' ** ''

Holothuria aphanes ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria arenacava ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria artensis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria fuscorubra ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria hilla ''Holothuria hilla'' is a species of sea cucumber in the subgenus ''Mertensiothuria'' of the genus '' Holothuria''. Some common names include the contractile sea cucumber, the sand sifting sea cucumber and the tigertail sea cucumber, and in Hawai ...

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Holothuria isuga ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria leucospilota'' ** '' Holothuria viridiaurantia'' * Subgenus ''Metriatyla'' ** '' Holothuria aculeata'' ** ''

Holothuria albiventer ''Holothuria (Metriatyla) albiventer'' is a species of sea cucumber in the genus '' Holothuria'', subspecies ''Metriatyla''. First described by Semper in 1868, the species is distributed in the Western Indian Ocean and the Red Sea The Red S ...

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Holothuria alex ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria brauni ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria conica ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria cyrielle ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria horrida'' ** ''

Holothuria keesingi ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria lessoni ''Holothuria lessoni'', the golden sandfish, is a species of sea cucumber Sea cucumbers are echinoderms from the class Holothuroidea (). They are marine animals with a leathery skin and an elongated body containing a single, branched gonad. ...

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Holothuria martensii ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' ** '' Holothuria scabra'' ** '' Holothuria submersa'' ** '' Holothuria tortonesei'' * Subgenus ''Microthele'' ** '' Holothuria fuscogilva'' ** ''

Holothuria fuscopunctata ''Holothuria fuscopunctata'', the elephant trunkfish, is a species of sea cucumber in the family Holothuriidae native to shallow water in the tropical Indo-Pacific. It is placed in the subgenus ''Microthele'', making its full name ''Holothuria ( ...

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Holothuria nobilis ''Holothuria'' (''Microthele'') ''nobilis'', the black teatfish, is a species of sea cucumber in the genus ''Holothuria''. The cucumber is found in the tropical waters of the Indo-pacific ocean. It was first described by Emil Selenka Emil Selen ...

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Holothuria whitmaei ''Holothuria (Microthele) whitmaei'', commonly known as the black teatfish, is a species of sea cucumber in the family Holothuriidae. The sea cucumber is distributed in the western Indian and Pacific oceans, with specimens being found off of Tan ...

'' * Subgenus ''Panningothuria'' ** '' Holothuria austrinabassa'' ** '' Holothuria forskali'' * Subgenus ''Platyperona'' ** ''

Holothuria crosnieri ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria difficilis'' ** ''

Holothuria excellens ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria insolita ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria parvula ''Holothuria parvula'', the golden sea cucumber, is a species of echinoderm in the class Holothuroidea. It was first described by Emil Selenka in 1867 and has since been placed in the subgenus ''Platyperona'', making its full scientific name ''Ho ...

'' ) ** '' Holothuria rowei'' ** '' Holothuria samoana'' ** '' Holothuria sanctori'' * Subgenus ''Roweothuria'' ** ''

Holothuria arguinensis ''Holothuria arguinensis'' is a species of sea cucumber in the family Holothuriidae and subgenus ''Roweothuria''. It is found in waters off the northeast Atlantic Ocean and Mediterranean Sea. According to some scholarly research, the species is a ...

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Holothuria poli ''Holothuria (Roweothuria) poli'', also known as the white spot cucumber, is a species of sea cucumber in the family Holothuridae and the subgenus ''Roweothuria.'' The species was first described by the Italian doctor and naturalist Stefano del ...

'' ** '' Holothuria vemae'' * Subgenus ''Selenkothuria'' ** ''

Holothuria bacilla ''Holothuria (Selenkothuria) bacilla'' is a species of sea cucumber Sea cucumbers are echinoderms from the class Holothuroidea (). They are marine animals with a leathery skin and an elongated body containing a single, branched gonad. Sea cuc ...

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Holothuria carere ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria erinacea'' ** '' Holothuria glaberrima'' ** ''

Holothuria lubrica ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria mactanensis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria moebii ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria parva ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria parvispinea ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria portovallartensis'' ** '' Holothuria sinica'' ** '' Holothuria theeli'' ** '' Holothuria vittalonga'' * Subgenus ''Semperothuria'' ** ''

Holothuria cinerascens ''Holothuria'' (''Semperothuria'') ''cinerascens'' is a species of sea cucumber in the family Holothuriidae. The sea cucumber is widely distributed in the Pacific and Indian ocean, being found from the Red Sea to Madagascar and from Japan to Austra ...

'' ** '' Holothuria flavomaculata'' ** ''

Holothuria granosa ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria imitans'' ** '' Holothuria languens'' ** ''

Holothuria roseomaculata ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria surinamensis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' * Subgenus ''Stauropora'' ** '' Holothuria aemula'' ** ''

Holothuria annulifera ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria bo'' ** '' Holothuria discrepans'' ** ''

Holothuria dofleinii ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria fuscocinerea'' ** ''

Holothuria hawaiiensis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria mitis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria modesta'' ** '' Holothuria olivacea'' ** '' Holothuria pervicax'' ** ''

Holothuria pluricuriosa ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' * Subgenus '' Stichothuria'' ** '' Holothuria coronopertusa'' * Subgenus ''Theelothuria'' ** '' Holothuria asperita'' ** ''

Holothuria cadelli ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria duoturriforma ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria foresti ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria hamata'' ** '' Holothuria imperator'' ** ''

Holothuria klunzingeri ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

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Holothuria kurti ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' ** '' Holothuria longicosta'' ** ''

Holothuria maculosa ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria michaelseni'' ** ''

Holothuria notabilis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria paraprinceps ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria princeps'' ** '' Holothuria pseudonotabilis'' ** '' Holothuria spinifera'' ** ''

Holothuria squamifera ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria turriscelsa ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' * Subgenus ''Thymiosycia'' ** ''

Holothuria altaturricula ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria arenicola'' ** '' Holothuria conusalba'' ** '' Holothuria gracilis'' ** '' Holothuria impatiens'' ** ''

Holothuria macroperona ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' ** '' Holothuria marginata'' ** ''

Holothuria milloti ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

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Holothuria minax ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' ** '' Holothuria remollescens'' ** '' Holothuria strigosa'' ** '' Holothuria thomasi'' ** ''

Holothuria truncata ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

'' ** '' Holothuria unicolor'' ** ''

Holothuria zihuatanensis ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' * Subgenus '' Vaneyothuria'' ** ''

Holothuria integra ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria lentiginosa'' ** '' Holothuria sinefibula'' ** ''

Holothuria suspecta ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' ** '' Holothuria unica'' ** ''

Holothuria zacae ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodied ...

'' * Subgenus '' incertae sedis'' (uncertain placement) ** '' Holothuria platei'' ** ''

Holothuria pyxoides ''Holothuria'' is the type genus of the marine animal family Holothuriidae, part of the class Holothuroidea, commonly known as sea cucumbers. Members of the genus are found in coastal waters in tropical and temperate regions. They are soft bodi ...

References

{{Taxonbar, from=Q1408044 Holothuriidae Holothuroidea genera