Holoplanktonic
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Holoplankton are organisms that are planktic (they live in the water column and cannot swim against a current) for their entire life cycle. Holoplankton can be contrasted with
meroplankton Meroplankton are a wide variety of aquatic organisms which have both planktonic and benthic stages in their life cycles. Much of the meroplankton consists of larval stages of larger organism. Meroplankton can be contrasted with holoplankton, whic ...
, which are planktic organisms that spend part of their life cycle in the
benthic zone The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning " ...
. Examples of holoplankton include some
diatom A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...
s, radiolarians, some
dinoflagellate The dinoflagellates (Greek δῖνος ''dinos'' "whirling" and Latin ''flagellum'' "whip, scourge") are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered algae. Dinoflagellates are ...
s, foraminifera,
amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far descr ...
s, krill,
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s, and salps, as well as some
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk species. Holoplankton dwell in the
pelagic zone The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
as opposed to the
benthic zone The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning " ...
. Holoplankton include both
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...
and zooplankton and vary in size. The most common plankton are protists.


Reproduction

Holoplankton have unique traits that make reproduction in the water column possible. Both sexual and asexual reproduction are used depending on the type of plankton. Some invertebrate holoplankton release sperm into the water column which are then taken up by the females for fertilization. Other species release both sperm and egg to increase the likelihood of fertilization. Environmental, mechanical, or chemical cues can all trigger this release. Diatoms are single celled phytoplankton that can occur as individuals or as long chains. They can reproduce sexually and asexually. Diatoms are important oxygen producers and are usually the first step in the food chain. Copepods are small holoplanktonic crustaceans that swim using their hind legs and antennae.


Defenses

Because of their small size and sluggish swimming abilities, holoplanktonic species have made certain specialized adaptations and in some cases are equipped with special defenses. Adaptations include flat bodies, lateral spines, oil droplets, floats filled with gases, sheaths made of gel-like substances, and ion replacement. Zooplankton have adapted by developing transparent bodies, bright colors, bad tastes and
cyclomorphosis Cyclomorphosis (also known as seasonal polyphenism) is the name given to the occurrence of cyclic or seasonal changes in the phenotype of an organism through successive generations. It occurs in small aquatic invertebrates that reproduce by parthen ...
(seasonal changes in body shape). When predators release a chemical in the water to signal zooplankton;
cyclomorphosis Cyclomorphosis (also known as seasonal polyphenism) is the name given to the occurrence of cyclic or seasonal changes in the phenotype of an organism through successive generations. It occurs in small aquatic invertebrates that reproduce by parthen ...
allows holoplankton to increase their spines and protective shields. Studies have shown that although small in size certain gelatinous zooplankton are rich in protein and lipid. "Many holoplankton seem to have very little visible defense mechanisms; therefore, it is hypothesized that a chemical defense may be possible. Pelagic cnidarians (jellyfish and related species) have
nematocyst A cnidocyte (also known as a cnidoblast or nematocyte) is an explosive cell containing one large secretory organelle called a cnidocyst (also known as a cnida () or nematocyst) that can deliver a sting to other organisms. The presence of this ce ...
s on their tentacles that eject a coiled microscopic thread very rapidly. These threads penetrate the surface of their target and release a series of complicated, biologically advanced venoms. Their stings can be very dangerous, due in part to the number of vital systems affected.


Sexual holoplankton

*
Copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...


See also

* Plankton *
Meroplankton Meroplankton are a wide variety of aquatic organisms which have both planktonic and benthic stages in their life cycles. Much of the meroplankton consists of larval stages of larger organism. Meroplankton can be contrasted with holoplankton, whic ...


Sources

Australian Museum Online


References

{{plankton Aquatic ecology Planktology