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The Himalayan shrike-babbler (''Pteruthius aeralatus ripleyi'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
subspecies found in the western Himalayas that belongs to the shrike-babbler group. The genus was once considered to be an aberrant
Old World babbler The Old World babblers or Timaliidae are a family of mostly Old World passerine birds. They are rather diverse in size and coloration, but are characterised by soft fluffy plumage. These are birds of tropical areas, with the greatest variety in S ...
and placed in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Timaliidae until molecular phylogenetic studies showed them to be closely related to the
vireo The vireos make up a family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. "Vireo" is a Latin word referring to a green migrato ...
s of the New World, leading to their addition in the family
Vireonidae The vireos make up a family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. "Vireo" is a Latin word referring to a green migratory bir ...
. Males and females have distinctive plumages, with the males being all black about with a cinnamon-rufous tertial patch and a distinctive white stripe running from behind the eye. The underside is whitish with some pinkish buff on the flanks. Females have a greyish head, lack the white stripe and have the upperparts and wings with greens, yellow and chestnut. The subspecies is part of a
cryptic species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
that was earlier considered as one species, white-browed shrike-babbler (''P. flaviscapis'' in the broad sense) with several subspecies.


Description

Adult males have a black head and a greyish back. A white superciliary stripe begins just above and behind the eye, extending back down the sides of the nape. The throat and underside are whitish. The inner edges and tips of the primaries are white, the tertiaries (the three inner secondaries) are uniformly orange-chestnut, unlike dark chestnut in ''P. a. validirostris''. The flanks have a hint of pale pink and grey. The grey on the back is lighter than the shade in ''validirostris''. The bill is black and hooked at the tip. The adult female is olive brown above with the wing coverts edged grey. The secondaries are edged with olive green and the outer three primaries are tipped in white on the inner edge. The tertiaries are uniformly chestnut. Both males and females are paler than ''P. a. validirostris''. The tail is black. The outer tail feathers are olive green and tipped in yellow with the inner webs black. The central tail feathers are olive green with a black tip. The underside is whitish buff as in the male but washed with grey from the middle of the abdomen to the vent.


Taxonomy

The taxonomic history of this subspecies is complex. It was originally described by Nicholas Aylward Vigors in 1831 who described the bird based on a specimen from Murree in Pakistan. He called it ''Lanius erythropterus'' and the subspecies was later moved out of the genus ''Lanius'' used only for true shrikes and placed in the genus ''Pteruthius'' as ''P. erythropterus''. In 1951, Koelz described a new subspecies from Nagaland under the name of ''Pteruthius erythropterus validirostris''. In 1960, Biswamoy Biswas noted that the name ''Lanius erythropterus'' was "preoccupied" or clashing with an earlier name used by Shaw in 1809 for a different species and he therefore suggested that the western Himalayan species should be renamed as a subspecies ''ripleyi'' after elevating Koelz's name for the eastern Himalayan form to the rank of a species. Temminck described a similar and related species from Java in 1835 as ''Allotrius flaviscapis'' and this too was moved to the genus ''Pteruthius''. The subspecies ''validirostris'', which then included ''ripleyi'', along with many other forms in the region were lumped as subspecies of ''Pteruthius flaviscapis'' by Ernst Mayr and R.A. Paynter in their taxonomic treatment in 1964. This was followed by most regional works such as the ''
Handbook of the Birds of India and Pakistan The ''Handbook of the Birds of India and Pakistan'' is the ''Masterpiece, magnum opus'' of Indian ornithologist Salim Ali, written along with S. Dillon Ripley. Appended to the title is the phrase "''together with those of Bangladesh, Nepal, Sikkim ...
'' by Salim Ali and Dillon S. Ripley. In 2008, a study of the ''Pteruthius'' group found that the relationships within the groups here were more complex than earlier thought and one of the findings was that the subspecies ''validirostris'' of the eastern Himalayas was more closely related to the forms found further to the east in Southeast Asia and far more distant to the form in the western Himalayas. This led to the elevation of the western form as ''Pteruthius ripleyi'' with ''validirostris'' being made a subspecies of ''Pteruthius aeralatus'' Blyth's shrike-babbler.


Distribution

The subspecies is found in the western Himalayas from northern Pakistan and extending east through India into central
Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...
and possibly further east. The eastern limits of the species are unclear but some evidence based on song differences suggests that they might occur as far east as Arunachal Pradesh. This might be negated if it is found that song variations exist within the eastern form ''Pteruthius aeralatus validirostris'' of Blyth's shrike-babbler.


Behaviour

Himalayan shrike-babblers are strictly arboreal and are seen foraging for insects mainly in the upper canopy. During the breeding season they are found in pairs but at other times several birds may be found, often in
mixed-species foraging flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. They also feed on berries, hopping along branches and sometimes hanging like a nuthatch. They sometimes sit still and will call often. The call being a series of loud ''kewkew kwekew'' repeated three or four times and the song from February to June transcribed as ''cha-chew, cha-ca-chip''. The nest is a hammock, like that of an oriole, built in a fork towards the tip of a branch high in the canopy of a tree. The clutch varies from two to four eggs which are pinkish white and speckled in purple brown, the spots merging to form a ring towards the broad end.


References

{{Taxonbar, from=Q10809501 Himalayan shrike-babbler Birds of the Himalayas Himalayan shrike-babbler Himalayan shrike-babbler