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''Heterocheila'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of acalyptrate
true flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
(Diptera). They are placed in their own family, Heterocheilidae, in the superfamily
Sciomyzoidea Sciomyzoidea is a superfamily of Acalyptratae flies. The families placed here include at least the following: * Coelopidae – seaweed flies * Dryomyzidae * Helcomyzidae * Helosciomyzidae * Heterocheilidae * Huttoninidae * Natalimyzi ...
. They are not widely familiar outside entomological circles, but the common name "half-bridge flies" has been associated with them.McAlpine, D. K. 1991. Relationships of the genus Heterocheila (Diptera: Sciomyzoidea) with description of a new family. Tijdschr. Ent. 134: 193-199. [1991.12.18
online
/ref> They are medium-sized flies occurring mainly in temperate regions on seashores of the Northern Hemisphere, where they and their larvae typically feed on stranded kelp in the wrack zone. In this, they resemble Coelopidae, kelp flies, which are members of a different family, though the same superfamily. The family Heterocheilidae was established by McAlpine in 1991. He distinguished it from other families to which ''Heterocheila'' had hitherto been referred at various times and by various authorities – Helcomyzidae, Dryomyzidae and
Coelopidae The Coelopidae or kelp flies are a family of Acalyptratae flies (order Diptera), they are sometimes also called seaweed flies, though both terms are used for a number of seashore Diptera. Fewer than 40 species occur worldwide. The family is fou ...
.


Description

For terms see
Morphology of Diptera Dipteran morphology differs in some significant ways from the broader morphology of insects. The Diptera is a very large and diverse order of mostly small to medium-sized insects. They have prominent compound eyes on a mobile head, and (at most) ...
The Heterocheilidae are medium to moderately large (body length 4.2-6.5 mm), fairly robust, brown flies Their postverical bristles are long and often parallel. Three fronto-orbital bristles are seen. The acrostichal bristles are in a single row with a larger prescutellar pair; the metepisternum is bare. The prosternum is triangular, and extended laterally to the propleuron as a narrow, precoxal bridge. The costa is unbroken and the crossveins are unmarked. The midregion of costa lacks anteroventral spines. The apical section of vein M is nearly straight, terminating posterior of wing apex. Crossvein Bm-Cu is almost aligned with vein CuA2. Vein CuA2+A1 reaches the wing margin.


Species

Two species have been described: * ''Heterocheila buccata'' Rondani, 1857 has a Palearctic distribution in North America and Eurasia. * ''Heterocheila hannai'' (Cole, 1921) has been reported from the NE USA coast of Alaska, Oregon, and Washington. The genus name ''Oedoparea'' Loew, 1862 is regarded as synonymous.Loew, H. 1862. Diptera. Ueber die europäischen Helomyzidae und die in Schlesien vorkommenden Arten derselben. Z. Ent. (Breslau) (1859) 13: 1-80


References


External links


Diptera.info
Images of ''Heterocheila buccata'' Sciomyzoidea genera Taxa named by Camillo Rondani {{Sciomyzoidea-stub