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''Hesperis matronalis'' is an herbaceous plant species in the family
Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some are shrubs. The leav ...
. It has numerous common names, including dame's rocket, damask-violet, dame's-violet, dames-wort, dame's gilliflower, night-scented gilliflower, queen's gilliflower, rogue's gilliflower, summer lilac, sweet rocket, mother-of-the-evening, Good & Plenties, and winter gilliflower. These plants are
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...
s or short-lived perennials, native to
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago ...
and cultivated in many other areas of the world for their attractive, spring-blooming flowers. In some of those areas, it has escaped from cultivation and become a weed species. The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
name ''Hesperis'' was probably given because the scent of the flowers becomes more conspicuous towards evening (''Hespera'' is the Greek word for evening).


Description

''Hesperis matronalis'' grows 100 cm or taller, with multiple upright, hairy stems. Typically, the first year of growth produces a mound of foliage, and flowering occurs the second year; the plants are normally
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...
s, but a number of races can be short-lived
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wid ...
s. The plants have showy blooms in early to mid spring. The leaves are alternately arranged on upright stems and lanceolate-shaped; they typically have very short or lack petioles and have toothed margins, but sometimes are entire and are widest at the base. The foliage has short hairs on the top and bottom surfaces that give the leaves a somewhat rough feel. The larger leaves are around 12 cm long and over 4 cm wide. In early spring, a thick mound of low-growing foliage is produced; during flowering the lower parts of the stems are generally unbranched and denuded of foliage and the top of the blooming plant might have a few branches that end in
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
s. The plentiful, fragrant flowers are produced in large, showy, terminal racemes that can be 30+ cm tall and elongate as the flowers of the inflorescence bloom. When stems have both flowers and fruits, the weight sometimes causes the stems to bend. Each flower is large (2 cm across), with four petals. Flower coloration varies, with different shades of lavender and purple most common, but white, pink, and even some flowers with mixed colors exist in cultivated forms. A few different double-flowered varieties also exist. The four petals are clawed and hairless. The flowers have six stamens in two groups, the four closest to the ovary are longer than the two oppositely positioned. Stigmas are two-lobed. The four
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s are erect and form a mock tube around the claws of the petals and are also colored similarly to the petals. Some plants may bloom until August, but warm weather greatly shortens the duration on each flower's blooming. Seeds are produced in thin fruits 5–14 cm long pods, containing two rows of seeds separated by a dimple. The fruit are
terete Terete is a term in botany used to describe a cross section that is circular, or like a distorted circle, with a single surface wrapping around it.Lichen Vocabulary, Lichens of North America Information, Sylvia and Stephen Sharnoff/ref> This is u ...
and open by way of
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
valves, constricted between the seeds like a pea pod. Seeds are oblong, 3–4 mm long and 1–1.5 mm wide. In North America, ''Hesperis matronalis'' is often confused with native ''
Phlox ''Phlox'' (; Greek φλόξ "flame"; plural "phlox" or "phloxes", Greek φλόγες ''phlóges'') is a genus of 67 species of perennial and annual plants in the family Polemoniaceae. They are found mostly in North America (one in Siberia) in di ...
'' species that also have similar large showy flower clusters. They can be distinguished from each other by foliage and flower differences: dame's rocket has alternately arranged leaves and four petals per flower, while phloxes have opposite leaves and five petals.


Cultivation

''Hesperis matronalis'' has been a cultivated species for a long time, and grows best in full sun to partial shade where soils are moist with good drainage. It is undemanding and self-seeds quickly, forming dense stands. Extensive monotypic stands of dame's rocket are visible from great distances; these dense collections of plants have the potential to crowd out native species when growing outside of cultivated areas. The successful spread of dame's rocket in North America is attributed to its prolific seed production and because the seeds are often included in prepackaged "wildflower seed" mixes sold for "naturalizing". The plants typically produce a low-lying rosette of foliage the first year; in subsequent years, blooming and seed production occurs in tandem throughout the blooming season. This species is commonly found in roadside ditches, dumps and in open woodland settings, where it is noticed when in bloom. Although it makes an attractive, hardy garden plant, it has been found to be ecologically invasive in North America. ''Hesperis matronalis'' is propagated by seeds, but desirable individuals, including the double-flowering forms, are propagated from cuttings or division of the clumps. Young leaves are high in vitamin C and can be eaten in salads and have a slightly bitter taste. Seeds can be sprouted and also eaten in salads.


Distribution

A garden escape, ''H. matronalis'' is found in many areas of Ireland, including Belfast. Dame's rocket was brought to North America in the 17th century and has since become naturalized there, now growing throughout most of the US and Canada. The US Department of Agriculture website has a map showing states and provinces in which the plant has been found. In Europe, it is host to the caterpillars of several
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprise ...
species, including the orange tip (''
Anthocharis cardamines ''Anthocharis cardamines'', the orange tip, is a butterfly in the family Pieridae, which contains about 1,100 species. ''A. cardamines'' is mainly found throughout Europe and temperate Asia (Palearctic) The males feature wings with a signature o ...
''), large white (''
Pieris brassicae ''Pieris brassicae'', the large white, also called cabbage butterfly, cabbage white, cabbage moth (erroneously), or in India the large cabbage white, is a butterfly in the family Pieridae. It is a close relative of the small white, ''Pieris ra ...
''), small white (''
Pieris rapae ''Pieris rapae'' is a small- to medium-sized butterfly species of the whites-and-yellows family Pieridae. It is known in Europe as the small white, in North America as the cabbage white or cabbage butterfly, on several continents as the small c ...
''), and
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s, such as '' Plutella porrectella''.


Weediness

It is considered an invasive species in some areas. Five U.S. states have placed legal restrictions on it: * In Colorado, it is classed as a noxious weed (B-list), with plans for eradication or management varying by area and year. * In Connecticut, it is classified as invasive and banned, making it illegal to move, sell, purchase, transplant, cultivate, or distribute the plant. * In Massachusetts, it is prohibited. * In New York, it is classified as invasive and eradication considered infeasible. * In Wisconsin, it is classed as restricted, i.e. an invasive species that is already widely established in the state, and causes, or has the potential to cause significant environmental or economic harm. In Alberta, Canada, it is considered a noxious weed, meaning its growth or spread needs to be prevented.


References


External links

* {{Taxonbar, from=Q1857886 Brassicaceae Garden plants Plants described in 1753 Taxa named by Carl Linnaeus