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A hesperidium (plural hesperidia) is a modified
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
with a tough, leathery rind.


Etymology

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
gave the name Hesperideæ to an order containing the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Citrus'', in allusion to the
golden apple The golden apple is an element that appears in various national and ethnic folk legends or fairy tales. Recurring themes depict a hero (for example Hercules or Făt-Frumos) retrieving the golden apples hidden or stolen by a monstrous antagonist. ...
s of the
Hesperides In Greek mythology, the Hesperides (; , ) are the nymphs of evening and golden light of sunsets, who were the "Daughters of the Evening" or "Nymphs of the West". They were also called the Atlantides () from their reputed father, the Titan Atlas ...
.


Development

The outer
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
wall becomes the thick spongy layer of the rind, while the inner ovary wall becomes very juicy with several seeds. The peel contains volatile oil glands in pits. The fleshy interior is composed of separate sections, called
carpels Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
, filled with fluid-filled vesicles that are specialized hair cells.


Uses

Oranges An orange is a fruit of various citrus species in the family Rutaceae (see list of plants known as orange); it primarily refers to ''Citrus'' × ''sinensis'', which is also called sweet orange, to distinguish it from the related ''Citrus × ...
,
lemon The lemon (''Citrus limon'') is a species of small evergreen trees in the flowering plant family Rutaceae, native to Asia, primarily Northeast India (Assam), Northern Myanmar or China. The tree's ellipsoidal yellow fruit is used for culin ...
s,
limes Limes may refer to: * the plural form of lime (disambiguation) * the Latin word for ''limit'' which refers to: ** Limes (Roman Empire) (Latin, singular; plural: ) is a modern term used primarily for the Germanic border defence or delimiting ...
, and
grapefruit The grapefruit (''Citrus'' × ''paradisi'') is a subtropical citrus tree known for its relatively large, sour to semi-sweet, somewhat bitter fruit. The interior flesh is segmented and varies in color from pale yellow to dark pink. Grapefruit is ...
are all common examples of hesperidia. Unlike most other
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
, the rind of cultivated hesperidia is generally not eaten with the fruit because it is tough and bitter. A common exception is the
kumquat Kumquats (; zh, 金桔), or cumquats in Australian English, are a group of small fruit-bearing trees in the flowering plant family Rutaceae. Their taxonomy is disputed. They were previously classified as forming the now-historical genus ''For ...
, which is consumed entirely. The outermost, pigmented layer of rind contains
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
s and is known as the ''flavedo''. When scraped off and used as a culinary ingredient it is called
zest Zest may refer to: Common usage * Zest (ingredient), the outer peel of a citrus fruit ** Zester, a tool for preparing zest ** Twist (cocktail garnish), a piece of zest * Zest (positive psychology), a component of character Brands * Zest (brand ...
. A confection called
succade Succade is the candied peel of any of the citrus species, especially from the citron or ''Citrus medica'' which is distinct with its extra-thick peel; in addition, the taste of the inner rind of the citron is less bitter than those of the other ...
can also be produced by candying the inner rind (known as
pith Pith, or medulla, is a tissue in the stems of vascular plants. Pith is composed of soft, spongy parenchyma cells, which in some cases can store starch. In eudicotyledons, pith is located in the center of the stem. In monocotyledons, it ext ...
or ''albedo'') of the
citron The citron (''Citrus medica''), historically cedrate, is a large fragrant citrus fruit with a thick rind. It is said to resemble a 'huge, rough lemon'. It is one of the original citrus fruits from which all other citrus types developed throu ...
or
lemon The lemon (''Citrus limon'') is a species of small evergreen trees in the flowering plant family Rutaceae, native to Asia, primarily Northeast India (Assam), Northern Myanmar or China. The tree's ellipsoidal yellow fruit is used for culin ...
.


See also

*
Fruit anatomy Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggre ...


References

*


External links

* Fruit morphology Rutaceae {{Rutaceae-stub