Hepatica In Lithuania
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''Hepatica'' (hepatica, liverleaf, or liverwort) is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of herbaceous
perennials A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
in the buttercup family, native to central and northern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
and eastern North America. Some botanists include ''Hepatica'' within a wider interpretation of ''
Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all continents except Australia, New Zealand an ...
''.


Description

Bisexual
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s with pink, purple, blue, or white
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s and three green bracts appear singly on hairy stems from late
winter Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultur ...
to
spring Spring(s) may refer to: Common uses * Spring (season) Spring, also known as springtime, is one of the four temperate seasons, succeeding winter and preceding summer. There are various technical definitions of spring, but local usage of ...
. Butterflies,
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s, bees,
flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced m ...
and
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s are known pollinators. The leaves are basal, leathery, and usually three-lobed, remaining over
winter Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultur ...
.


Taxonomy

''Hepatica'' was described by the English botanist
Philip Miller Philip Miller FRS (1691 – 18 December 1771) was an English botanist and gardener of Scottish descent. Miller was chief gardener at the Chelsea Physic Garden for nearly 50 years from 1722, and wrote the highly popular ''The Gardeners Dicti ...
in 1754. It was proposed as a subgenus of ''
Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all continents except Australia, New Zealand an ...
'' in 1836, but later segregated into genus ''Hepatica''.


Taxa

,
Kew Kew () is a district in the London Borough of Richmond upon Thames. Its population at the 2011 census was 11,436. Kew is the location of the Royal Botanic Gardens ("Kew Gardens"), now a World Heritage Site, which includes Kew Palace. Kew is a ...
's Plants of the World Online (POWO) accepts 7 
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and one
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
in the genus ''Hepatica'': One infraspecific
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
is also recognized by POWO: * ''Hepatica nobilis'' var. ''japonica'' ** Synonym: ''Hepatica asiatica'' ** Synonym: ''Hepatica insularis'' ''Hepatica'' can be divided into two
series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...
with respect to leaf edge:


Series ''Triloba''

The leaves of the series ''Triloba'' Ulbr.Ulbrich, O.E.: Über die systematische Gliederung und geographische Verbreitung der Gattung Anemone L. Bot. Jahrb. Syst. (1905) 37: 172 - 257, 38: 257 - 334.
Tamura Tamura (usually written 田村), a Japanese placename and family name, may refer to: In places: *Tamura, Fukushima, a city in Japan *Tamura District, Fukushima, in Japan * Tamura Station, in Nagahama, Japan People with the surname Tamura: * Tamura ...
:Tamura, M.: Morphology, ecology and phylogeny of the Ranunculaceae” VII. Science reports of South College, North College of Osaka University, Japan 16:21-43, 1968. are three-lobed with a smooth leaf edge. * ''Hepatica acutiloba'':
Eastern Canada Eastern Canada (also the Eastern provinces or the East) is generally considered to be the region of Canada south of the Hudson Bay/ Strait and east of Manitoba, consisting of the following provinces (from east to west): Newfoundland and Labrado ...
,
Midwestern United States The Midwestern United States, also referred to as the Midwest or the American Midwest, is one of four census regions of the United States Census Bureau (also known as "Region 2"). It occupies the northern central part of the United States. I ...
,
Eastern United States The Eastern United States, commonly referred to as the American East, Eastern America, or simply the East, is the region of the United States to the east of the Mississippi River. In some cases the term may refer to a smaller area or the East C ...
* ''Hepatica americana'': Central Canada, Eastern Canada, Midwestern United States, Eastern United States * ''Hepatica maxima'': endemic to Ulleung-do island (South Korea) * ''Hepatica nobilis'': ** ''Hepatica nobilis'' var. ''nobilis'':
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
to
Western Siberia Western Siberia or West Siberia (russian: Западная Сибирь, Zapadnaya Sibir'; kk, Батыс Сібір) is a part of the larger region of Siberia that is mostly located in the Russian Federation. It lies between the Ural region an ...
** ''Hepatica nobilis'' var. ''japonica'':
Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...
, North China,
Central China Central China () is a geographical and a loosely defined cultural region that includes the provinces of Henan, Hubei and Hunan. Jiangxi is sometimes also regarded to be part of this region. Central China is now officially part of South Centra ...
,
East China East China () is a geographical and a loosely defined cultural region that covers the eastern coastal area of China. A concept abolished in 1978, for economical purposes the region was defined from 1949 to 1961 by the Chinese Central Govern ...
,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
, and Japan (
Honshu , historically called , is the largest and most populous island of Japan. It is located south of Hokkaidō across the Tsugaru Strait, north of Shikoku across the Inland Sea, and northeast of Kyūshū across the Kanmon Straits. The island se ...
,
Shikoku is the smallest of the four main islands of Japan. It is long and between wide. It has a population of 3.8 million (, 3.1%). It is south of Honshu and northeast of Kyushu. Shikoku's ancient names include ''Iyo-no-futana-shima'' (), '' ...
, north of Kyushu island)


Series ''Angulosa''

The leaves of series ''Angulosa'' (Ulbr.) Tamura are three- to five-lobed with a
crenate A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
leaf edge. * ''Hepatica falconeri'' — Mountain forests of Central Asia; India: Northwest Himalayas (
Himachal Pradesh Himachal Pradesh (; ; "Snow-laden Mountain Province") is a state in the northern part of India. Situated in the Western Himalayas, it is one of the thirteen mountain states and is characterized by an extreme landscape featuring several peaks ...
, Jammu and Kashmir); Northwest China ( Tienshan); Kyrgyzstan and Tajikistan ( Pamir-Alai); North Pakistan, Kazakhstan (North Tienshan) * ''Hepatica henryi'': North Central China,
South Central China South Central China, South-Central China or Central-South China ( zh, c = 中南, p = Zhōngnán, l = Central-South), is a region of the People's Republic of China defined by State Council that includes the provinces of Guangdong, Hainan, He ...
(western
Hubei Hubei (; ; alternately Hupeh) is a landlocked province of the People's Republic of China, and is part of the Central China region. The name of the province means "north of the lake", referring to its position north of Dongting Lake. The ...
, northern
Hunan Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi ...
,
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
,
Shaanxi Shaanxi (alternatively Shensi, see § Name) is a landlocked province of China. Officially part of Northwest China, it borders the province-level divisions of Shanxi (NE, E), Henan (E), Hubei (SE), Chongqing (S), Sichuan (SW), Gansu (W), N ...
) * ''Hepatica × media'':
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central, Eastern, and Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, Serbia to the southwest, Moldova to the east, and ...
* ''Hepatica transsilvanica'': Romania ( Carpathian Mountains,
Transylvania Transylvania ( ro, Ardeal or ; hu, Erdély; german: Siebenbürgen) is a historical and cultural region in Central Europe, encompassing central Romania. To the east and south its natural border is the Carpathian Mountains, and to the west the Ap ...
)


Etymology

The word ''hepatica'' derives from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
, from 'liver', because its three-lobed leaf was thought to resemble the human
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it ...
.


Distribution

Plants of genus ''Hepatica'' are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, and North America. * Europe: Albania, Austria, the Baltic states, Belarus, Bulgaria, Corsica, Czechoslovakia, Denmark, European Russia, Finland, France, Germany, Greece, Hungary, Italy, Norway, Poland, Romania, Spain, Sweden, Switzerland, Ukraine, Yugoslavia * Central Asia: Kazakhstan, Kyrgyzstan, Tajikistan, Western Siberia * Eastern Asia: North China,
South Central China South Central China, South-Central China or Central-South China ( zh, c = 中南, p = Zhōngnán, l = Central-South), is a region of the People's Republic of China defined by State Council that includes the provinces of Guangdong, Hainan, He ...
,
East China East China () is a geographical and a loosely defined cultural region that covers the eastern coastal area of China. A concept abolished in 1978, for economical purposes the region was defined from 1949 to 1961 by the Chinese Central Govern ...
, Japan, Korea, Manchuria, Primorsky Krai * South Asia: Pakistan, Western Himalaya * Canada: Manitoba, New Brunswick, Nova Scotia, Ontario, Québec * United States: Alabama, Arkansas, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Kentucky, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Vermont, Virginia, West Virginia, Wisconsin Plants of the genus have been introduced to Belgium.


Cultivation

''Hepatica'' cultivation has been popular in Japan since the 18th century (mid-
Edo period The or is the period between 1603 and 1867 in the history of Japan, when Japan was under the rule of the Tokugawa shogunate and the country's 300 regional '' daimyo''. Emerging from the chaos of the Sengoku period, the Edo period was characte ...
), where flowers with doubled petals and a range of colour patterns have been developed. Noted for its tolerance of alkaline
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
-derived soils, ''Hepatica'' may grow in a wide range of conditions; it can be found either in deeply shaded deciduous (especially beech)
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (se ...
and scrub or
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
in full sun. ''Hepatica'' will also grow in both sandy and clay-rich substrates, being associated with
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
. Moist soil and winter snowfall are required; ''Hepatica'' is tolerant of winter snow cover, but less so of dry frost. Propagation is done by seeds or by dividing vigorous clumps in spring. However, seedlings take several years to reach bloom size, and divided plants are slow to thicken.


Uses

''Hepatica'' was once used as a
medicinal herb Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including defense and protection ag ...
. Owing to the
doctrine of signatures The doctrine of signatures, dating from the time of Dioscorides and Galen, states that herbs resembling various parts of the body can be used by herbalists to treat ailments of those body parts. A theological justification, as stated by botanist ...
, the plant was once thought to be an effective treatment for liver disorders. Although poisonous in large doses, the leaves and flowers may be used as an
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian pla ...
, as a
demulcent A demulcent (derived from the la, demulcere "caress") is a mucilaginous or oleaginous preparation that forms a soothing protective film over a mucous membrane, relieving minor pain and inflammation of the membrane. However, they generally help fo ...
for slow-healing injuries, and as a
diuretic A diuretic () is any substance that promotes diuresis, the increased production of urine. This includes forced diuresis. A diuretic tablet is sometimes colloquially called a water tablet. There are several categories of diuretics. All diuretics i ...
.


References


External links

* * * {{Taxonbar, from=Q157505 Ranunculaceae genera Medicinal plants of Asia Medicinal plants of Europe Medicinal plants of North America Taxa named by Philip Miller