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The hen harrier (''Circus cyaneus'') is a
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
. It breeds in
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
. The term "hen harrier" refers to its former habit of preying on free-ranging fowl. It migrates to more southerly areas in winter. Eurasian birds move to southern Europe and southern temperate Asia. In the mildest regions, such as France and
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
, hen harriers may be present all year, but the higher ground is largely deserted in winter. The
northern harrier The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost United States, USA. The northern ...
was formerly considered to be a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the hen harrier.


Taxonomy

In 1758 the English naturalist George Edwards included an illustration and a description of the hen harrier in the first volume of his ''Gleanings of Natural History''. He used the English name "The blue hawk". Edwards based his hand-coloured etching on a bird that had been shot near London. When in 1766 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' for the twelfth edition, he placed the hen harrier with the falcons and eagles in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Falco''. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Falco cyaneus'' and cited Edwards' work. The hen harrier is now placed in the genus ''
Circus A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicy ...
'' that was introduced by the French naturalist
Bernard Germain de Lacépède Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French natural history, naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's g ...
in 1799. The genus name ''Circus'' is derived from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''kirkos'', referring to a bird of prey named for its circling flight (''kirkos'', "circle"). The specific epithet ''cyaneus'' is from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
and means "dark blue". The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised. The hen harrier was formerly considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
with the
northern harrier The northern harrier (''Circus hudsonius''), also known as the marsh hawk or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost United States, USA. The northern ...
.


Description

The hen harrier is long with a wingspan. It resembles other harriers in having distinct male and female plumages. The sexes also differ in weight, with males weighing , with an average of , and females weighing , with an average of . Among standard measurements, the wing chord is , the
tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
is and the tarsus is . It is relatively long winged and long tailed. The male is mainly grey above and white below except for the upper breast, which is grey like the upperparts, and the rump, which is white; the wings are grey with black wingtips. The female is brown above with white upper tail coverts, hence females, and the similar juveniles, are often called "ringtails". Their underparts are buff with brown barring. Juveniles resemble females but with less distinct barring, dark brown secondaries dark brown and less-barred belly. The female gives a whistled ''piih-eh'' when receiving food from the male, and her alarm call is ''chit-it-it-it-it-et-it''. The male calls ''chek-chek-chek'', with a more bouncing ''chuk-uk-uk-uk'' during his display flight.


Behaviour

This medium-sized raptor breeds on
moorland Moorland or moor is a type of Habitat (ecology), habitat found in upland (geology), upland areas in temperate grasslands, savannas, and shrublands and the biomes of montane grasslands and shrublands, characterised by low-growing vegetation on So ...
,
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and musk ...
s,
prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
s,
farm A farm (also called an agricultural holding) is an area of land that is devoted primarily to agricultural processes with the primary objective of producing food and other crops; it is the basic facility in food production. The name is used fo ...
land coastal prairies,
marsh In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...
es,
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and other assorted open areas. A male will maintain a territory averaging , though male territories have ranged from . These are one of the few raptorial birds known to practice
polygyny Polygyny () is a form of polygamy entailing the marriage of a man to several women. The term polygyny is from Neoclassical Greek πολυγυνία (); . Incidence Polygyny is more widespread in Africa than in any other continent. Some scholar ...
– one male mates with several females. Up to five females have been known to mate with one male in a season. A supplementary feeding experiment on the Orkney islands showed that rates of polygyny were influenced by food levels; males provided with extra food had more breeding females than 'control' males that received no extra food. The nest is constructed on the ground or on a mound of dirt or vegetation. Nests are made of sticks and are lined inside with grass and leaves. Four to eight (exceptionally 2 to 10) whitish eggs are laid. The eggs measure approximately . The eggs are incubated mostly by the female for 31 to 32 days. When incubating eggs, the female sits on the nest while the male hunts and brings food to her and the chicks. The male will help feed chicks after they hatch, but does not usually watch them for a greater period of time than around 5 minutes. The male usually passes off food to the female, which she then feeds to the young, although later the female will capture food and simply drop into the nest for her nestlings to eat. The chicks fledge at around 36 days old, though breeding maturity is not reached until 2 years in females and 3 years in males. In winter, the hen harrier is a bird of open country, and will then roost communally, often with merlins and
marsh harrier The marsh harriers are bird of prey, birds of prey of the harrier (bird), harrier subfamily. They are medium-sized Bird of prey, raptors and the largest and broadest-winged harriers. Most of them are associated with marshland and dense reedbeds. ...
s. There is now an accepted record of transatlantic vagrancy by the northern harrier, with a juvenile being recorded in Scilly,
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
from October 1982 to June 1983.


Hunting behaviour

This is a typical harrier, which hunts on long wings held in a shallow V in its low flight during which the bird closely hugs the contours of the land below it. Northern or hen harriers hunt primarily small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s, as do most harriers. Up to 95% of the diet comprises small mammals. However,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s are hunted with some regularity as well, especially by males. Preferred avian prey include
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
s of open country (i.e. sparrows,
lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
s, pipits), small shorebirds and the young of
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...
and galliforms. Supplementing the diet occasionally are
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s (especially
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s),
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s and
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s (especially
orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – gras ...
ns). The species has been observed to hunt
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s if these are available. Larger prey, such as
rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s and adult
duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
s are taken sometimes and harriers have been known to subdue these by drowning them in water. Harriers hunt by surprising prey while flying low to the ground in open areas, as they drift low over fields and moors. The harriers circle an area several times listening and looking for prey. Harriers use hearing regularly to find prey, as they have exceptionally good hearing for diurnal raptors, this being the function of their
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
-like facial disc. This harrier tends to be a very vocal bird while it glides over its hunting ground.


Mortality and competition

Little information is available on the longevity in hen harriers. The longest-lived known bird is 16 years and 5 months. However, adults rarely live beyond 8 years. Early mortality is mainly due to predation. Predators of eggs and nestlings include
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...
s,
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
s,
badger Badgers are medium-sized short-legged omnivores in the superfamily Musteloidea. Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by their squat bodies and adaptions for fossorial activity rather than by the ...
s,
fox Foxes are small-to-medium-sized omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull; upright, triangular ears; a pointed, slightly upturned snout; and a long, bushy tail ("brush"). Twelve species ...
es, crows and ravens,
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s and
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s. Both parents attack potential predators with alarm calls and striking with talons.
Short-eared owl The short-eared owl (''Asio flammeus'') is a widespread grassland species in the family Strigidae. Owls belonging to genus ''Asio'' are known as the eared owls, as they have tufts of feathers resembling mammalian ears. These "ear" tufts may or ...
s are natural competitors of this species, preferring the same prey and habitat, and having a similar geographic distribution. Occasionally, both harriers and short-eared owls will harass each other until the victim drops its prey and it can be stolen, a practice known as
kleptoparasitism Kleptoparasitism (originally spelt clepto-parasitism, meaning "parasitism by theft") is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct f ...
. Most often, the harriers are the aggressors, stealing prey from the owls.


Status

This species has a wide geographical distribution and, although there is evidence of a population decline, it is not thought to be approaching the thresholds for the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
criterion of population decline (i.e., a decline of more than 30% in ten years or three generations). It is therefore classified as "least concern". However, in Britain and Ireland, hen harrier populations are in a critical state, mainly due to habitat loss and illegal shooting on grouse moors. There is sufficient suitable habitat to support over 300 breeding pairs of hen harriers in England alone, yet only 34 successful nests were recorded in 2022. Furthermore, many of the young birds do not survive beyond their first year, before the onset of sexual maturity at the age of two. Most UK Hen Harriers are found in Scotland, but even there the population declined by 27% between 2004 and 2016.


Relationship with humans

In some parts of Europe people believed that seeing a hen harrier perched on a house was a sign that three people would die. Unlike many raptors, hen harriers have historically been looked upon favorably by farmers because they eat predators of quail eggs and mice that damage crops. Harriers are sometimes called "good hawks" because they do not pose a threat to poultry as some hawks do.


Forestry and hen harriers

The hen harrier is a bird of open habitats such as heather moorland and extensive agricultural land. However, much of its range, particularly in Ireland and parts of western Britain, has been (and continues to be) afforested, mainly with non-native conifers such as
Sitka spruce ''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to just over tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth- ...
(''Picea sitchensis'') from North America. Hen harriers nest and forage in commercial forestry plantations before the canopy closes (typically at between 9 and 12 years old), but do not make much use of thicket and later growth stages, which typically comprise between and of the commercial growth cycle. Where afforestation replaces habitats previously used by hen harriers they will therefore tend to reduce overall habitat availability. However, where afforestation takes place in areas that were previously underutilised by hen harriers, it may increase the value of such areas to the species in the long-term. Areas dominated by forestry may remain suitable to hen harriers provided that a mosaic of age classes is maintained within the forest so that areas of young, pre-thicket forest are always available.


References


External links


The Langholm Moor Demonstration Project

Field Guide Page on Flickr


(in Persian)
Ageing and sexing (PDF; 4.3 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
*
RSPB Skydancer website
* *
Save The Skydancer page
on BirdWatchIreland.ie {{DEFAULTSORT:harrier, hen
hen harrier The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Palearctic, Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl. It bird migration, migrates to more southerly areas in winter. Eurasian ...
Birds of prey of Eurasia
hen harrier The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Palearctic, Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl. It bird migration, migrates to more southerly areas in winter. Eurasian ...
hen harrier The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Palearctic, Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl. It bird migration, migrates to more southerly areas in winter. Eurasian ...