Hemorrhois Hippocrepis
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The horseshoe whip snake (''Hemorrhois hippocrepis'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Colubridae Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
. The species is native to southwestern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and northern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
.


Description

Adults of ''H. hippocrepis'' may attain a total length (including tail) of 1.5 m (5 feet). Its body is slender, and its head is wider than its neck. The eye is large, with a round pupil, and with a row of small scales below it. The smooth
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are arranged in 25-29 rows, and the
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
number 220–258.
Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, it has a series of large spots which are either blackish or dark brown edged with black. There is a series of alternating smaller dark spots on each side. The lighter ground color between the spots may be yellowish, olive, or reddish. The dark spots are closely spaced, giving the appearance of a dark snake with a light pattern resembling a chain or a series of X's. There is a light horseshoe-shaped mark on the neck and back of head.


Geographic range

''H. hippocrepis'' is found in
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
and
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
in North Africa, and in southern and central
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
, southern, eastern and central
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Gibraltar ) , anthem = " God Save the King" , song = " Gibraltar Anthem" , image_map = Gibraltar location in Europe.svg , map_alt = Location of Gibraltar in Europe , map_caption = United Kingdom shown in pale green , mapsize = , image_map2 = Gib ...
, southern
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
and
Pantelleria Island Pantelleria (; Sicilian: ''Pantiddirìa'', Maltese: ''Pantellerija'' or ''Qawsra''), the ancient Cossyra or Cossura, is an Italian island and comune in the Strait of Sicily in the Mediterranean Sea, southwest of Sicily and east of the Tunis ...
in Europe. In the island locations, it may have been introduced. Since the early 2000s it has been reported from
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
of
Mallorca Mallorca, or Majorca, is the largest island in the Balearic Islands, which are part of Spain and located in the Mediterranean. The capital of the island, Palma, is also the capital of the autonomous community of the Balearic Islands. The Bal ...
,
Ibiza Ibiza (natively and officially in ca, Eivissa, ) is a Spanish island in the Mediterranean Sea off the eastern coast of the Iberian Peninsula. It is from the city of Valencia. It is the third largest of the Balearic Islands, in Spain. Its l ...
and
Formentera Formentera (, ) is the smallest and most southerly island of the Pityusic Islands group (comprising Ibiza and Formentera, as well as various small islets), which belongs to the Balearic Islands autonomous community (Spain). It covers an area of ...
. It could have been introduced there by way of old olive trees imported from mainland Spain. The species is thriving there and becomes larger than on the mainland.Ellie Shechet, August 24, 2020.
A Plague on Ibiza


Habitat

The natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of ''H. hippocrepis'' are Mediterranean-type shrubby vegetation, rocky areas, rocky shores, sandy shores,
arable land Arable land (from the la, arabilis, "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for the ...
, pastureland,
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s, rural gardens, and
urban area An urban area, built-up area or urban agglomeration is a human settlement with a high population density and infrastructure of built environment. Urban areas are created through urbanization and are categorized by urban morphology as cities, t ...
s.


Reproduction

''H. hippocrepis'' is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
.


Subspecies

Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. *''Hemorrhois hippocrepis hippocrepis'' *''Hemorrhois hippocrepis nigrescens'' Pantelleria ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a genus other than ''Hemorrhois''.


Conservation status

The horseshoe whip snake is assessed as being of "
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
" by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
in its
Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
. Its population trend is thought to be steady, and it is able to adapt to modified habitats. Threats it faces include being run over by traffic, poisoned by agricultural chemicals and being captured for use by local snake charmers.


See also

*
List of reptiles of Italy The Italian reptile fauna totals 58 species (including introduced and naturalised species). They are listed here in three systematic groups (Sauria, Serpentes, and Testudines) in alphabetical order by scientific name. Sauria (lizards) *'' Algyroi ...


References


Further reading

* Cattaneo A (1985). "''Il colubro ferro di cavallo dell'Isola di Pantelleria: ''Coluber hippocrepis nigrescens subsp. nova'' (Reptilia Squamata Colubridae)''". ''Atti della Società italiana di Scienze naturali del Museo civico di Storia naturale di Milano'' 126 (3-4): 165–184. (''Coluber hippocrepis nigrescens'', new subspecies). (in Italian, with an abstract in English). * Linnaeus C (1758). ''Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, diferentiis, synonymis, locis. Tomus I. Editio Decima, Reformata.'' Stockholm: L. Salvius. 824 pp. (''Coluber hippocrepis'', new species, p. 226). (in Latin). * Utiger U, Helfenberger N, Schätti B, Schmidt C, Ruf M, Ziswiler V (2002). "Molecular Systematics and Phylogeny of Old World Ratsnakes, ''Elaphe'' Auct., and Related Genera (Reptilia, Squamata, Colubridae)". ''Russian Journal of Herpetology'' 9 (2): 105–124. (''Hemorrhois hippocrepis''). (in English). {{Taxonbar, from=Q540927 Hemorrhois Reptiles of Europe Reptiles of North Africa Fauna of the Iberian Peninsula Reptiles described in 1758 Taxa named by Carl Linnaeus Taxonomy articles created by Polbot