The butterfly sculpin (''Hemilepidotus papillon'') is a species of fish in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Agonidae Agonidae is a family of small, bottom-dwelling, cold-water marine fish. Common names for members of this family include poachers, Irish lords, sea ravens, alligatorfishes, starsnouts, hooknoses, and rockheads. They are notable for having elongate ...

. It is found in the North Pacific Ocean.

Taxonomy

The butterfly sculpin was first formally described in 1880 as ''Melletes papilio'' by the American

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...

Tarleton Hoffman Bean Tarleton Hoffman Bean (October 8, 1846 – December 28, 1916) was an American ichthyologist. Biography and education Tarleton Hoffman Bean was born to George Bean and Mary Smith Bean in Bainbridge, Pennsylvania, on October 8, 1846. He attended ...

with its type locality given as Saint Paul Island in the

Pribilof Islands The Pribilof Islands (formerly the Northern Fur Seal Islands; ale, Amiq, russian: Острова Прибылова, Ostrova Pribylova) are a group of four volcanic islands off the coast of mainland Alaska, in the Bering Sea, about north of ...

in the

Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...

off

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

. Bean proposed the monospecific genus ''Melletes' fo the butterfly sculpin but later workers have placed it in the genus ''

Hemilepidotus ''Hemilepidotus'', the Irish lords, is a genus of ray-finned fishes belonging to the family Agonidae, the poachers and sea ravens. These fishes are found in northern Pacific, northern Atlantic and the Arctic Oceans. Species There are currently s ...

''. The specific name ''papilio'' means "butterfly".

Description

The butterfly sculpin is reddish brown, yellow and white in color, with a metallic gold sheen. There are four blackish bars on the upper flanks which extend onto the dorsal fin, these bands vary in their definition. This fish has between 11 and 13 spines and 19 or 20 soft rays in its dorsal fins while there are no spines and between 16 and 18 soft rays in its

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

.The scales in the ventral scale row are less than half the size of the scales in the dorsal scale row. The membrane of the first, spiny

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

is deeply notched. There are four spines on the preoperculum with the fourt spine being simple. The maximum published

fork length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...

is , and maximum weight of .

Distribution

The butterfly sculpin lives in

benthopelagic The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...

marine environments of the northern

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

and the

Sea of Okhotsk The Sea of Okhotsk ( rus, Охо́тское мо́ре, Ohótskoye móre ; ja, オホーツク海, Ohōtsuku-kai) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands ...

, in sandy or silty flat areas. It is usually found at depths of and water temperature from −0.9 to 10.7 °C.

Biology

Butterfly sculpins are predators feeding on gammarid amphipods, crabs,

ostracods Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typical ...

and small fish such as juvenile

Alaska pollock The Alaska pollock or walleye pollock (''Gadus chalcogrammus'') is a marine fish species of the cod genus ''Gadus'' and family Gadidae. It is a semi-pelagic Shoaling and schooling, schooling fish widely distributed in the North Pacific Ocean, N ...

. They are

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

laying benthic eggs which hatch into pelagic larvae which settle on the substrate as juveniles at a minimum length of .

References

{{Taxonbar, from=Q2313899 Butterfly sculpin Fish described in 1880 Taxa named by Tarleton Hoffman Bean