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''Hemiancistrus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of suckermouth armored catfishes. These species are native to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. The
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of this genus is complex and unclear, and major work has to be done. Many of these fish are popular
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
fish.


Taxonomy

''Hemiancistrus'' is a genus within the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
Ancistrini Ancistrini is a tribe of catfishes of the family Loricariidae. Most are restricted to tropical and subtropical South America, but there are also several genus (''Ancistrus'', '' Chaetostoma'', ''Hemiancistrus'' and ''Lasiancistrus'') in southern ...
of the subfamily
Hypostominae The Hypostominae are a subfamily of catfishes of the family Loricariidae. Most members are restricted to tropical and subtropical South America, but there are also several species (in genera ''Ancistrus'', '' Chaetostoma'', ''Lasiancistrus'', '' ...
. This genus has long been used as a "dump" for Loricariid species; fish with unclear relationships have been classified as members of this genus. As such, this taxon is not
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
. At this point, many undescribed species remain. This genus and the closely related ''
Peckoltia ''Peckoltia'' is a genus of small South American Loricariidae, armored suckermouth catfishes. Many of these fish are popular aquarium fish. Taxonomy ''Peckoltia'' is a Basal (phylogenetics), basal genus within the tribe Ancistrini of the subfami ...
'' may be
synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
, as neither genus is supported by
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
. ''Hemiancistrus'' species differ from other members of the ''
Panaque The genus ''Panaque'' contains a small number of small to medium-sized South American suckermouth armoured catfishes that are notable for being among the very few vertebrates that feed extensively on wood. In addition, algae and aufwuchs are a ...
''
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
lacking the synapomorphies of the other genera and having the
dentaries In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
meeting at an angle greater than 120°; in ''Peckoltia'' species, the dentaries meet at less than 90° Generally, ''Peckoltia'' are considered to be those that have dorsal saddles and bands in the fins, while ''Heminancistrus'' have spots and uniform coloration. What species belong to this genus has been unclear. Many newer species have been tentatively assigned to ''Hemiancistrus''. Although the recently described ''H. micrommatos'' and ''H. spinosissimus'' have been considered
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of ''H. spilomma'' by Armbruster, they have more recently been considered valid. '' H. micrommatos'', '' H. spinosissimus'' and '' H. spilomma'' are currently placed in the genus ''
Ancistomus ''Ancistomus'' is a genus of suckermouth armored catfishes found in shallow waters in rapidly flowing rivers in the southeastern Amazon basin in Brazil. Species There are currently 5 recognized species in this genus: * '' Ancistomus feldberga ...
''. ''
Hemiancistrus medians ''Hemiancistrus medians'' is a species of catfish in the family Loricariidae. It is native to South America, where it occurs in the Maroni basin. This species is usually found in rapids with a strong current and a substrate of large, shelter-pro ...
'' is larger than most species that were left in ''Hemiancistrus'', has well-developed keels (only present elsewhere in '' ‘H.’ landoni''), and a different body shape. The remainder of the taxa that do not have established genera that they can be placed in will be recognized as species groups in ''‘Hemiancistrus’'' in single quotes until they can be examined further.


Species

There are currently 12 recognized species in this genus: * ''
Hemiancistrus cerrado ''Hemiancistrus cerrado'' is a species of catfish in the family Loricariidae. It is native to South America, where it occurs in the Araguaia River basin in the Tocantins River drainage in Brazil. It is usually found in rocky riffles in second-o ...
'' L. S. de Souza, M. R. S. de Melo, Chamon & Armbruster, 2008 * ''
Hemiancistrus chlorostictus ''Hemiancistrus chlorostictus'' is a species of catfish in the family Loricariidae. It is native to South America, where it occurs in the Uruguay River basin, including the Passo Fundo River, in Brazil Brazil ( pt, Brasil; ), officially t ...
'' A. R. Cardoso & L. R. Malabarba, 1999 * '' Hemiancistrus fuliginosus'' A. R. Cardoso & L. R. Malabarba, 1999 * '' Hemiancistrus furtivus'' Provenzano & Barriga, 2017 * '' Hemiancistrus guahiborum'' Werneke, Armbruster, Lujan & Taphorn, 2005 * ''
Hemiancistrus landoni ''Hemiancistrus landoni'' is a species of catfish in the family Loricariidae. It is native to South America, where it occurs in the Guayas River basin in Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), o ...
'' C. H. Eigenmann, 1916 * ''
Hemiancistrus medians ''Hemiancistrus medians'' is a species of catfish in the family Loricariidae. It is native to South America, where it occurs in the Maroni basin. This species is usually found in rapids with a strong current and a substrate of large, shelter-pro ...
'' Kner, 1854 * '' Hemiancistrus megalopteryx'' A. R. Cardoso, 2004 * '' Hemiancistrus meizospilos'' A. R. Cardoso & J. F. P. da Silva, 2004 * '' Hemiancistrus punctulatus'' A. R. Cardoso & L. R. Malabarba, 1999 * '' Hemiancistrus subviridis'' Werneke, Sabaj Pérez, Lujan & Armbruster, 2005 * '' Hemiancistrus votouro'' A. R. Cardoso & J. F. P. da Silva, 2004


Distribution and habitat

''Hemiancistrus'' species are mainly found in tropical and subtropical South America east of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
, but there is also species (''‘H.’ landoni'') in Pacific coast drainages of
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
. The genus exhibits a large distribution area ranging from the Panama, in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, to southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. Many ''Hemiancistrus'' originate from the
Guyanas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * G ...
, the
Negro In the English language, ''negro'' is a term historically used to denote persons considered to be of Black African heritage. The word ''negro'' means the color black in both Spanish and in Portuguese, where English took it from. The term can be ...
and
Orinoco The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wor ...
and the southern
Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technology c ...
ian tributaries. These fish prefer flowing water habitats of medium to large rivers.


Appearance and anatomy

''Hemiancistrus'' are members of the family Loricariidae, the armored suckermouth catfishes. As such, they have armor plating on their body instead of
scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number w ...
. Also, they have a
suckermouth In fish, a suckermouth is a ventrally-oriented (inferior) mouth adapted for grazing on algae and small organisms that grow on submerged objects. All Loricariidae possess a suckermouth as do the cypriniform algae eaters of the genus ''Gyrinocheilus' ...
which they use to cling to rocks in their habitat. They have the characteristic Loricariid
omega iris The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are ...
as well. Like many other catfish, ''Hemiancistrus'' have strong pectoral and dorsal fin spines that can be locked outwards as a defense. ''Hemiancistrus'' species are rather small to medium-sized Loricariid species. The largest species of the genus, '' ‘H.’ megalopteryx'', reaches about 29 centimetres (12 in). These fish also tend to be spotted or uniform in coloration.


In the aquarium

''Hemiancistrus'' species are popular
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
fish. Many species are relatively small and attractively colored. '' ‘Hemiancistrus’ subviridis'' is one such example, a bright green fish with yellow spots. However, '' ‘H.’ guahiborum'' is only rarely imported; despite being common in its natural habitat, it is less attractive than ''‘H.’ subviridis'', which occurs in the same area.


References

{{Taxonbar, from=Q1758739 Ancistrini Fish of South America Fish of the Amazon basin Freshwater fish genera Catfish genera Taxa named by Pieter Bleeker