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Helicinidae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of small tropical
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
s which have an operculum. They are
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
operculate
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Helicinoidea Helicinoidea is a taxonomic superfamily of land snails that have an operculum. In other words, they are terrestrial operculate gastropod mollusks. They are in the superfamily Neritoidea and are quite closely related to the marine and freshwa ...
. These snails are not at all closely related to the air-breathing land snails, despite a superficial similarity of the shells. The name "Helicinidae" immediately reminds most people of "
Helicidae Helicidae is a large, diverse family (biology), family of small to large, air-breathing land snails, sometimes called the "typical snails." A number of species in this family are valued as food items, including ''Cornu aspersum'' (formerly ''Heli ...
", the most popular group of land snails including e.g. Helix pomatia, but the important two letters "ni" only from the Latin diminutive of the latter name alluding to superficial similarities of otherwise completely different things.


Distribution

These snails are found in tropical and subtropical areas but they have an odd distribution: they are restricted to the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
islands and some
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
and Pacific islands, as well as the edge of the
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
n and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
n continents. Some species are found in the southern United States, from Louisiana to Florida; others in Central and South America.


Taxonomy

Helicinidae belongs to superfamily
Helicinoidea Helicinoidea is a taxonomic superfamily of land snails that have an operculum. In other words, they are terrestrial operculate gastropod mollusks. They are in the superfamily Neritoidea and are quite closely related to the marine and freshwa ...
according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). This family consists of six following subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): * Helicininae Férussac, 1822 - synonym: Olygyridae Gray, 1847; Bourcierinae Paetel, 1890 * Ceratodiscinae Pilsbry, 1927 * † Dimorphoptychiinae Wenz, 1938 * Hendersoniinae H. B. Baker, 1926 * Stoastomatinae C. B. Adams, 1849 *
Vianinae Vianinae is a subfamily of tropical land snail, a terrestrial gastropod mollusk in the family Helicinidae. Species According to the World Register of Marine Species, the following genera are accepted within ''Vianinae'': * '' Calidviana'' ...
H. B. Baker, 1922


Genera

Genera within the family Helicinidae include: Subfamily Helicininae * ''
Alcadia ''Alcadia'' is a genus of tropical and subtropical land snails with an operculum, terrestrial gastropod mollusks in the family Helicinidae. Species Species within the genus ''Alcadia'' include: * ''Alcadia affinis'' C.B. Adams, 1846 * '' A ...
'' Gray, 1840 * '' Aphanoconia'' A. J. Wagner, 1905 * ''
Bourciera ''Bourciera'' is a genus of neotropical terrestrial gastropod mollusks or land snails in the family Helicinidae. All species in the genus have an operculum. In 1907, an attempt was made to rename the genus ''Pseudhelicina'', sometimes corr ...
'' Pfeiffer, 1852 * '' Ceochasma'' F. G. Thompson, 1968 * '' Ceratopoma'' Möllendorff, 1893 * ''
Emoda ''Emoda'' is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Helicinidae Helicinidae is a family of small tropical land snails which have an operculum. They are terrestrial operculate gastropod mollu ...
'' Adams, 1856 * '' Glyptemoda'' Clench & Aguayo, 1950 * ''
Helicina ''Helicina'' is a genus of tropical and subtropical land snails with an operculum, terrestrial gastropod mollusks. This is the type genus of its family (Helicinidae) and subfamily ( Helicinae), as well as the superfamily Helicinoidea. This ...
'' Lamarck, 1799 - type genus of the family Helicinidae * '' Hemipoma'' Wagner, 1905 * '' Kosmetopoma'' A. J. Wagner, 1905 * '' Negopenia'' Iredale, 1941 * '' Nesiocina'' Richling & Bouchet, 2013 * '' Ogasawarana'' Wagner, 1905 * '' Olygyra'' Say, 1818: synonym of ''
Helicina ''Helicina'' is a genus of tropical and subtropical land snails with an operculum, terrestrial gastropod mollusks. This is the type genus of its family (Helicinidae) and subfamily ( Helicinae), as well as the superfamily Helicinoidea. This ...
'' Lamarck, 1799 (original rank) * '' Orobophana'' Wagner, 1905'' Phil. Trans. R. Soc. Lond.'' B (1998): 353, 347. * '' Palaeohelicina'' A. J. Wagner, 1905 * '' Pecoviana'' Iredale, 1941 * '' Pleuropoma'' Moellendorff, 1893 * '' Schasicheila'' Shuttleworth, 1852 * '' Stoastomops'' Baker, 1924 * '' Sturanya'' Wagner, 1905 * '' Sulfurina'' Möllendorff, 1893 subfamily Ceratodiscinae * '' Ceratodiscus'' Simpson & Henderson, 1901 - type genus of the subfamily Ceratodiscinae Subfamily † Dimorphoptychiinae * † '' Dimorphoptychia'' Sandberger, 1871 - type genus of the subfamily Dimorphoptychiinae Subfamily Hendersoniinae * '' Hendersonia'' A. J. Wagner, 1905 - type genus of the subfamily Hendersoniinae * '' Waldemaria'' Wagner, 1905 Subfamily Stoastomatinae * '' Stoastoma'' C. B. Adams, 1849 - type genus of the subfamily Stoastomatinae Subfamily
Vianinae Vianinae is a subfamily of tropical land snail, a terrestrial gastropod mollusk in the family Helicinidae. Species According to the World Register of Marine Species, the following genera are accepted within ''Vianinae'': * '' Calidviana'' ...
* '' Calidviana'' Baker, 1954 * ''
Calybium The calybium and the cupule make up the accessory fruit of flowering plants in the family Fagaceae. These two parts derive from different flower components. The cupule holds and protects the fruit during its growth and maturation. In some gen ...
'' Morelet, 1891 * '' Eutrochatella'' Fischer, 1885 * '' Geophorus'' Fischer, 1885 * '' Heudeia'' Crosse, 1885 * '' Lucidella'' Swainson, 1840 * '' Pseudotrochatella'' Nevill, 1881 * '' Pyrgodomus'' Fischer & Crosse, 1893 * '' Viana'' H. Adams & A. Adams, 1856 - type genus of the subfamily Vianinae Subfamily ? * '' Priotrochatella'' Fischer, 1893 * '' Semitrochatella'' Aguayo & Jaume, 1958 * '' Sturanyella'' Pilsbry & Cooke, 1934Mollusc Specialist Group (1996)
''Sturanyella carolinarum''
In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.1. . Downloaded on 17 June 2011.
* '' Troschelviana'' Baker, 1922 * ''
Ustronia ''Ustronia'' is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Helicinidae Helicinidae is a family of small tropical land snails which have an operculum. They are terrestrial operculate gastropod moll ...
'' Kobelt, 1908"Mollusca"
. Diversidad Biológica Cubana, accessed 23 March 2011.


References


Further reading

* Richling I. (2001). "New species of Helicinidae from Costa Rica (Gastropoda: Neritopsina)". ''Schriften zur Malakozoologie'' 17: 1-8. Cismar. * Richling I. (2004). "Classification of the Helicinidae: Review of Morphological Characteristics Based on a Revision of the Costa Rican Species and Application to the Arrangement of the Central American Mainland Taxa (Mollusca: Gastropoda: Neritopsina)". '' Malacologica'' 45(2): 195-440. * Richling I. (2004). "Coloration in Helicinidae (Mollusca: Gastropoda: Neritopsina)". ''
Malacologia ''Malacologia'' is a peer-reviewed scientific journal in the field of malacology, the study of mollusks. The journal publishes articles in the fields of molluscan systematics, ecology, population ecology, genetics, molecular genetics, evolu ...
'' 46(1): 217-224. * Richling I. (2005). "Biographical remarks on Dr.
Antoni Józef Wagner Antoni is a Catalan language, Catalan, Polish language, Polish, and Slovene language, Slovene given name and a surname used in the eastern part of Spain, Poland and Slovenia. As a Catalan given name it is a variant of the male names Anton (given nam ...
and the collection of Helicinidae in the Museum and Institute of Zoology in Warszawa". ''Folia Malacologica'' 13(4): 197-206. * Richling I. & Glaubrecht M. (2008). "The types of Neotropical Helicinidae (Mollusca, Gastropoda, Neritopsina) in the Malacological Collection of the Museum für Naturkunde Berlin: an annotated catalogue, with emphasis on Cuban land snails". ''
Zoosystematics and Evolution ''Zoosystematics and Evolution'' is a peer-reviewed open access scientific journal covering zoological systematics and evolution. It was established in 1898 as ''Mitteilungen aus der Zoologischen Sammlung des Museums für Naturkunde in Berlin'' ...
'' 84(2): 265-310. . * Richling I. (2009). "The Radiation of the Helicinidae in New Caledonia (Mollusca: Gastropoda: Neritopsina) including zoogeographical considerations". In: Grandcolas P. (ed.). "Zoologia Neocaledonica 7. Biodiversity studies in New Caledonia". ''Mémoires du Muséum national d'Histoire naturelle'' 198: 247-372. Paris
abstract
species specific reading: * Richling I., Franke S., Fernánndez V. A. & Sigaretta V., S. (2007). "New data on the micro-land snails ''Eutrochatella'' (''Microviana'') ''spinopoma'' Aguayo 1943 and ''Eutrochatella'' (''Microviana'') ''holguinensis'' Aguayo 1932 (Neritopsina: Helicinidae) in the province of
Holguín Holguín () is a municipality and city in Cuba, and the capital of Province of Holguín. After Havana, Santiago de Cuba, and Camagüey, it is the fourth largest city in Cuba. History Before Columbus, the Taino people settled in huts made fro ...
, eastern Cuba". ''Schriften zur Malakozoologie'' 23: 19-24. Cismar.


External links

* * Richling I
Forschung an Helicinidae
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by Ira Richling {{Taxonbar, from=Q2617976 Gastropod families