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''Hebius'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of snakes in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Colubridae Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
..


Geographic range

The genus ''Hebius'' is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
.Guo et al.


Taxonomy

All of the member
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the genus ''Hebius'' were formerly placed in the genus ''
Amphiesma Amphiesma is a genus of snakes in the family Colubridae. They are found across Asia. Species * '' Amphiesma stolatum'' (Linnaeus, 1758) – buff striped keelback * ''Amphiesma monticola ''Amphiesma monticola'', also known as the Wynad keel ...
'', but in 2014
Guo "Guo", written in Chinese: 郭, is one of the most common Chinese surnames and means "the wall that surrounds a city" in Chinese. It can also be transliterated into English as Cok, Gou, Quo, Quach, Quek, Que, Keh, Kuo, Kwo, Kuoch, Kok, Koc, ...
et al. placed most species of ''Amphiesma'' in the genus ''Hebius''. They placed other species in the genus ''
Herpetoreas ''Herpetoreas'' is a genus of snakes in the subfamily Natricinae of the family Colubridae. The genus is endemic to South Asia. Species The genus ''Herpetoreas'' contains the following 7 species, which are recognized as being valid. *'' Herpeto ...
'', leaving ''Amphiesma'' a
monotypic taxon In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
containing only the species ''
Amphiesma stolatum The buff striped keelback (''Amphiesma stolatum'') is a species of nonvenomous colubrid snake found across Asia. It is a typically nonaggressive snake that feeds on frogs and toads. It belongs to the subfamily Natricinae, and is closely related ...
''.


Species

The following 46 species are recognized as being valid. *''
Hebius andreae Andrea's keelback (''Hebius andreae'') is a species of nonvenomous colubrid snake endemic to Vietnam. Etymology The specific name, ''andreae'', is in honor of Andrea Ziegler, wife of German herpetologist Thomas Ziegler.Beolens, Bo; Watkins, Mi ...
'' ( Ziegler & Quyet, 2006) – Andrea’s keelback *''
Hebius annamensis The Annam keelback (''Hebius annamensis'') is a species of snake in the family Colubridae. It is found in Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southea ...
'' Bourret, 1934 – Annam keelback *''
Hebius arquus ''Hebius arquus'' is a species of snake of the family Colubridae. The snake is found in Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It cons ...
'' (
David David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...
& G. Vogel, 2010)
*''
Hebius atemporalis ''Hebius atemporalis'' is a species of snake of the family Colubridae. It is also known as the mountain keelback or Tonkin keelback. Geographic range The snake is found in northern Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), offic ...
'' (Bourret, 1934) – Tonkin keelback *''
Hebius beddomei The Nilgiri keelback (''Hebius beddomei''), also known commonly as Beddome's keelback, is a species of snake found in the Western Ghats in India. The species is named after Richard Henry Beddome, 1830–1911, British army officer and natural ...
'' ( Günther, 1864) – Nilgiri keelback, Beddome’s keelback *''
Hebius bitaeniatus ''Hebius bitaeniatus'', commonly known as the two-striped keelback or Kutkai keelback, is a species of snake of the family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by ...
'' (
Wall A wall is a structure and a surface that defines an area; carries a load; provides security, shelter, or soundproofing; or, is decorative. There are many kinds of walls, including: * Walls in buildings that form a fundamental part of the supe ...
, 1925)
– Kutkai keelback *''
Hebius boulengeri ''Hebius boulengeri'' is a species of snake in the family Colubridae. The species is native to East Asia and Southeast Asia. It is also known commonly as the Tai-yong keelback or Boulenger's keelback. Etymology The specific name, ''boulengeri' ...
'' ( Gressitt, 1937) – Tai-yong keelback, Boulenger’s keelback *''
Hebius celebicus ''Hebius celebicus'' the Sulawesi keelback, is a species of snake of the family Colubridae. The snake is found in Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian a ...
'' ( W. Peters &
Doria Doria or Dória may refer to: People Surname * Doria (family), a prominent Genoese family ** Andrea Doria (1466–1560), Genoese admiral ** Ansaldo Doria, 12th century Genoese statesman and commander ** Brancaleone Doria (died c. 1409?), husband ...
, 1878)
– Sulawesi keelback *''
Hebius chapaensis The Vietnam water snake or Chapa flat-nosed snake (''Hebius chapaensis'') is a species of colubrid snake. It is found in northern Vietnam and Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. ...
'' Bourret, 1934 – Vietnam water snake *''
Hebius clerki ''Hebius clerki'' the Yunnan keelback, is a species of snake of the family Colubridae. The snake is found in India, Myanmar, China, and Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic ...
'' (Wall, 1925) – Yunnan keelback *'' Hebius concelarus'' (
Malnate Malnate is a town and '' comune'' located in the province of Varese, in the Lombardy region of northern Italy. It lies in a mountainous region approximately north of Milan, in the foothills of the Alps The Alps () ; german: Alpen ; it, A ...
, 1963)
*'' Hebius craspedogaster'' ( Boulenger, 1899) – Kuatun keelback *''
Hebius deschauenseei ''Hebius deschauenseei'', commonly known as the northern keelback or Deschauensee's keelback, is a species of nonvenomous snake in the subfamily Natricinae of the family Colubridae. The species is endemic to Asia. Etymology The specific nam ...
'' (
Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) **List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) Plac ...
, 1934)
– northern keelback, Deschauensee’s keelback *'' Hebius flavifrons'' (Boulenger, 1887) – Sabah keelback *'' Hebius frenatus'' ( Dunn, 1923) – bridled keelback *'' Hebius groundwateri'' ( M.A. Smith, 1922) – Groundwater's keelback *'' Hebius igneus'' David, G. Vogel, T.Q. Nguyen, Orlov, Pauwels, Teynié & Ziegler, 2021 *'' Hebius inas'' (
Laidlaw Laidlaw (), organized as Laidlaw International, Inc. (with corporate headquarters in Naperville, Illinois) was the largest provider of intercity bus services, contract public transit and paratransit, and contract school bus service in both t ...
, 1901)
– Malayan mountain keelback *'' Hebius ishigakiensis'' (Malnate & Munsterman, 1960) – Yaeyama keelback *'' Hebius johannis'' (Boulenger, 1908) – Johann's keelback *'' Hebius kerinciensis'' (David & Das, 2003) *'' Hebius khasiensis'' (Boulenger, 1890) – Khasi Hills keelback, Khasi keelback *'' Hebius lacrima'' Purkayastha & David, 2019 – crying keelback *''
Hebius leucomystax The white-lipped keelback (''Hebius leucomystax'') is a species of Snake venom, nonvenomous Natricinae, natricine snake found in central Vietnam, Cambodia, and Laos. Etymology The Specific name (zoology), specific name, ''leucomystax'', is Gre ...
'' (David,
Bain Bain may refer to: People * Bain (surname), origin and list of people with the surname * Bain of Tulloch, Scottish family * Bain Stewart, Australian film producer, husband of Leah Purcell * Saint Bain (died c. 711 AD), Bishop of Thérouanne, Ab ...
, T.Q. Nguyen, Orlov, G. Vogel, Thanh & Ziegler, 2007)
– white-lipped keelback *'' Hebius metusia'' (
Inger Inger may refer to: People * Inger (given name), a list of people * Inger, the main character of Hans Christian Andersen's fairy tale ''The Girl Who Trod on a Loaf'' * Robert F. Inger (1920–2019), American herpetologist * Stella Inger, America ...
, E. Zhao, Shaffer & G. Wu, 1990)
– Wa Shan keelback *'' Hebius miyajimae'' (
Maki Maki may refer to: People *Mäki, a Finnish surname (includes a list of people with the name) *Maki (name), a Japanese given name and surname (includes a list of people with the name) Places *Maki, Ravar, Kerman Province, Iran *Maki, Rigan, Ke ...
, 1931)
– Maki's keelback *'' Hebius modestus'' (Günther, 1875) – modest keelback *'' Hebius nicobariensis'' ( Sclater, 1891) – Nicobar Island keelback *'' Hebius nigriventer'' (Wall, 1925) *'' Hebius octolineatus'' (Boulenger, 1904) – eight-lined keelback *'' Hebius optatus'' ( Hu & E. Zhao, 1966) – Mount Omei keelback *'' Hebius parallelus'' (Boulenger, 1890) – Yunnan keelback *'' Hebius petersii'' (Boulenger, 1893) – Peters's keelback *'' Hebius popei'' (
Schmidt Schmidt may refer to: * Schmidt (surname), including list of people with the surname * Schmidt (singer) (born 1990), German pop and jazz singer * Schmidt (lunar crater), a small lunar impact crater * Schmidt (Martian crater), a List of craters on ...
, 1925)
– Pope's keelback *'' Hebius pryeri'' (Boulenger, 1887) – Pryer's keelback *'' Hebius sanguineus'' ( Smedley, 1931) – Cameron Highlands keelback *'' Hebius sangzhiensis'' Zhou et al., 2019 – Sangzhi keelback *'' Hebius sarasinorum'' (Boulenger, 1896) – Sarasin’s keelback *'' Hebius sarawacensis'' (Günther, 1872) – Sarawak keelback *'' Hebius sauteri'' (Boulenger, 1909) – Kosempo keelback *'' Hebius septemlineatus'' (Schmidt, 1925) – Tengchong keelback *'' Hebius taronensis'' (M.A. Smith, 1940) – Kachin keelback *'' Hebius terrakarenorum'' Hauser, Smits, & David, 2022 *'' Hebius venningi'' (Wall, 1910) – Chin Hills keelback *'' Hebius vibakari'' ( H. Boie, 1826) – Japanese keelback *'' Hebius viperinus'' ( Schenkel, 1901) *'' Hebius weixiensis''
Hou Hou or HOU may refer to: * -hou, a place-name element * Hou (surname) * Hou (currency) (Chinese: ), a unit of currency in Greater China * Hou (Odder Municipality), a town in Denmark * Hou (title) (Chinese: ), a title in ancient China * Denglong (m ...
, Yuan, Wei, G. Zhao, G. Liu, Y. Wu, Shen, Chen,
Guo "Guo", written in Chinese: 郭, is one of the most common Chinese surnames and means "the wall that surrounds a city" in Chinese. It can also be transliterated into English as Cok, Gou, Quo, Quach, Quek, Que, Keh, Kuo, Kwo, Kuoch, Kok, Koc, ...
& Che, 2021
– Weixi keelback *'' Hebius yanbianensis'' Q. Liu, Zhong,
Wang Wang may refer to: Names * Wang (surname) (王), a common Chinese surname * Wāng (汪), a less common Chinese surname * Titles in Chinese nobility * A title in Korean nobility * A title in Mongolian nobility Places * Wang River in Thailand ...
, Y. Liu & Guo, 2018)
– Yanbian keelback ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Hebius''.


References


Further reading

* Guo, Peng; Zhu, Fei; Liu, Qin; Zhang, Liang; Li, Jian X.; Huang, Yu Y.; Pyron, R. Alexander (2014). "A taxonomic revision of the Asian keelback snakes, genus ''Amphiesma'' (Serpentes: Colubridae: Natricinae), with description of a new species". ''Zootaxa'' 3873 (4): 425–440. * Thompson JC (1913). "Contributions to the Anatomy of the Ophidia". ''Proceedings of the Zoological Society of London'' 1913: 414–425. (''Hebius'', new genus, p. 424). {{Taxonbar, from=Q18616085
Hebius ''Hebius'' is a genus of snakes in the family Colubridae.. Geographic range The genus ''Hebius'' is endemic to Asia.Guo et al. Taxonomy All of the member species of the genus ''Hebius'' were formerly placed in the genus ''Amphiesma'', but in 20 ...
Snake genera