Headwater Chub
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The headwater chub (''Gila nigra'') is a species of fish in the family
Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...
. It is found in Arizona and New Mexico.


Description

The body of the headwater chub is thick and chunky to streamlined but not markedly attenuate. The maximum size of the male fish is about 50 cm females are about 10–18 cm total length. The coloration of the fish is dark olive-gray or brown above, with a silver side, and white below. The longitudinal stripes are often diffuse, and are rarely with dark dorsal-lateral blotches. The caudal peduncle is not pencil-like, its length is less than its head length; the fins are small to moderate in size, sometimes convex, and are rarely curved, with inter-radial membranes of fins variously pigmented. Scales are developed and cover the entire body, with the basal radii variable. There are 73-83 lateral scales, with usually 8 dorsal and anal fin rays, and rarely 7 or 9 fin rays.Minckley, W.L. and B.D. DeMarais, 2000. ''Taxonomy of Chubs (Teleostei, Cyprinidae, Genus Gila) in the American Southwest with comments on conservation''. ''Copeia'', 2000(1), pp. 251-256. Similar species include the humpback chub (''Gila cypha'') and bonytail chub (''G. elegans''), however, these fish have extremely slender caudal peduncles, smaller eyes, angle along anal fin base continuing above the caudal fin. Large individuals have a hump on their nape, and a depressed head which is absent on the headwater chub. ''Gila nigra'' are somewhat trout-like in appearance, except they lack an adipose fin. Not surprisingly, they are morphologically intermediate between the
roundtail chub The roundtail chub (''Gila robusta'') is a cyprinid fish in the genus '' Gila'', of southwestern North America. It is native to the Colorado River drainage basin, including the Gila River and other tributaries, and in several other rivers. It i ...
(''Gila robusta'') and the Gila chub (''Gila intermedia'').


Range

Headwater chubs are endemic to the
Gila River The Gila River (; O'odham ima Keli Akimel or simply Akimel, Quechan: Haa Siʼil, Maricopa language: Xiil) is a tributary of the Colorado River flowing through New Mexico and Arizona in the United States. The river drains an arid watershed of n ...
basin of Arizona and New Mexico where they occupy the middle and headwater reaches of middle-sized streams. Populations have been recognized from the mainstream Gila River (above confluence with Mangus Creek) in New Mexico, this includes West, Middle and East forks of the Gila River, along with the San Carlos River (a tributary to the Gila). They are also identified from Ash Creek (tributary to San Carlos River),
Tonto Creek Tonto Creek is a stream located in the Mogollon Rim area of the state of Arizona on the north edge of the Tonto National Forest. The closest town, Payson, Arizona, Payson, is away. Tonto Creek is a stream that flows year round, starting just ...
(tributary to the Salt River), and Spring Creek, (tributary of Tonto Creek). In the Verde River system, they inhabit Upper Fossil Creek (above the diversion dam), East Verde River and Deadman Creek.Bestgen, K.R. and D.L. Probst. 1989. ''Distribution, status and notes on the ecology of Gila robusta in the Gila River drainage'', New Mexico. ''The Southwestern Naturalist'' 34(3): pp 402-412.Animal Abstract:''Gila nigra''
. Compiled by the Heritage Data Management System, Arizona Game and Fish Department, 2003.


Habitat

Adult headwater chub occupy cool to warm water in mid- to headwater stretches of mid-sized streams of the Gila River basin. They are associated with deep, near shore pools adjacent to swift riffles and runs, and near obstructions. Cover consists of root wads, boulders, undercut banks, submerged organic debris, or deep water. In Fossil Creek, they were found in water more than 1.8 m deep with velocities under 0.10 meters per second. Substrates they are associated with include gravel, small boulders, and large in-stream objects. Preferred water temperature ranges of 20-27 °C with a minimum temperature around 7 °C. Juveniles are associated with shallow, low velocity habitat with overhead cover. In Fossil Creek, headwater chub seem to select depths between 0.9-1.5 m and velocities of 0.15 meters per second and are found over sand substrate.Voeltz, J.B. 2002. ''Roundtail Chub (“Gila robusta”) status survey of the Lower Colorado River Basin''. AGFD Nongame and Endangered Wildlife Program Technical Report #186, Arizona Game and Fish Department. Phoenix, Arizona. pp. 221. The headwater chub are associated with substrates including gravel, small boulders, and large in-stream objects. The preferred water temperature ranges of the headwater chub are 20-27 °C with a minimum temperature around 7 °C. Juveniles are associated with shallow, low velocity habitat with overhead cover. In Fossil Creek, they seem to select depths between 0.9-1.5 m and velocities of 0.15 mps and are found over sand substrate. The headwater chub life span is 8–10 years. They grow rapidly but growth is dependent on water temperature. The maximum size of the fish is about 50 cm.


Population trends

As with many native fish, reductions in range and numbers are likely the result of habitat loss, as well as competition with, and predation by, non-native fish species.


Management factors

Activities that are known to be detrimental to populations should be avoided, including dewatering of habitats through re-routing stream water, stream impoundment, channelization, domestic livestock grazing, timber harvesting, mining, road construction, polluting, and stocking non-native fish. *Threats: aquifer pumping; stream diversion; reduction in stream flows; predation by and competition with nonnative fishes. *Management needs: watershed and stream flow protection; research to determine mechanisms of disappearance; ameliorate effects of deleterious nonnative fishes.


Protective measures taken

This species is currently being considered for T & E listing by the US Fish and Wildlife Service.


References


External links


List of Arizona Native Fishes

Arizona Desert Museum
{{DEFAULTSORT:Headwater chub Endangered fauna of the United States Chubs (fish) Gila (fish) Fauna of the Southwestern United States Freshwater fish of the United States Taxa named by Edward Drinker Cope Fish described in 1875