Harpa Conference Center Reykjavik City Center Hiticeland

''Harpa'', common name the "harp snails", is a genus of large predatory sea snails, marine gastropod mollusks in the family Harpidae.Gofas, S.; Bouchet, P. (2015). Harpa Röding, 1798. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=205853 on 2016-02-28

''Harpa'' is the type genus of the family Harpidae.

Description

The shell has an ovate-oblong shape. It is more or less inflated, generally pretty thin, enamelled, and provided with parallel, longitudinal, inclined and acute ribs. ; The body whorl is much larger than all the others together. The spire is slightly elevated. The aperture is large, oval, dilated, strongly emarginated inferiorly, and without siphonal canal. The outer lip is bordered by the last rib. The columella is smooth, simple, nearly straight and pointed at the base.

The animal has a flattened head, which supports a pair of pretty long, thick, and conical tentacles, with a small protuberance at their base, internally, where the eyes are situated. the mouth is simple, surrounded by a muscular margin, and furnished with a small, slender and pointed trunk. The organ of excitement is elongated, cylindrical, situated on the right side. The locomotive organ is very large, very broad at the anterior part, which is ear-shaped, and distinguished by a deep emargination upon each side. The posterior extremity is caducous, and destitute of an operculum.

The fleshy part of this mollusk is very strong, and very large. Its foot is enormous, thick, and extended considerably out of the shell. It cannot be wholly contained within the aperture, before which, by contracting itself, it forms a margin.

The foot is as if divided into two portions. The anterior broader, arcuated, ear-shaped, with a marginal furrow, and joined to the posterior part by a kind of neck. This latter, more extended, is somewhat oval, pointed, and slightly inflated above, without any appearance of operculum. When the animal is violently disturbed, it breaks off the posterior extremity of its foot, in order to withdraw itself more completely within its shell. In consequence of this an operculum would be useless to it, for it would be liable to be carried away by the rupture of the foot. Therefore, it is not possessed.

All the external parts of the animal are strongly colored with spots and plates of a brownish red, intermingled with other yellowish spots. The middle portion is frequently crossed by a brown band.

The respiratory tube is long, rather large, continued to and terminating in a large pulmonary cavity, the use of which is to assist respiration. Upon one side of this cavity are two pectinated branchiae, one large, and the other small. Upon the opposite side are situated, in female specimens, the rectum and the uterus ; and in the male a deferential canal and a penis. On this same side are fixed, at the upper part of the cavity, the mucous follicles, composed of seven or eight transverse plates. The tongue is small, slender, pointed, fleshy, without any trace of a ribbon of horn. It is contained in an incurved sheath, and rarely protruded from it. The stomach is very narrow, and does not differ in size from the rest of the intestinal canal. The rectum is pretty large, terminated by an anus slightly narrowed and pointed. The liver, which is voluminous, forms a great part of the convoluted portion, and extends almost throughout the spire. The heart and the auricle are very much developed, contained in a pericardium, and situated at the base of the branchiae. The cerebral ganglion is broad and flattened ; it sends out numerous nervous filaments which ramify over the whole body. The penis of the male is considerable and situated upon the same side.
Kiener (1840). General species and iconography of recent shells
: comprising the Massena Museum, the collection of Lamarck, the collection of the Museum of Natural History, and the recent discoveries of travellers; Boston :W.D. Ticknor,1837

This genus are voracious nocturnal predators of benthic crustaceans on sandy bottoms, capable of eating crabs as large as themselves.

Distribution

This marine species is circumtropical, except the western Atlantic Ocean. It also occurs off Australia ( Northern Territory, Queensland, Western Australia).

Species

Species in the genus ''Harpa'' include:

* '' Harpa amouretta'' Röding, 1798
* '' Harpa articularis'' Lamarck, 1822
* '' Harpa cabriti'' P. Fischer, 1860
* '' Harpa costata'' (Linnaeus, 1758)
* '' Harpa crenata'' Swainson, 1822
* '' Harpa davidis'' Röding, 1798
* '' Harpa doris'' Röding, 1798
* '' Harpa goodwini'' Rehder, 1993
* ''Harpa gracilis'' Broderip & G. B. Sowerby I, 1829
* ''Harpa harpa'' (Linnaeus, 1758)
* ''Harpa kajiyamai'' Habe, 1970
* '' Harpa kolaceki'' T. Cossignani, 2011
* ''Harpa lorenzi'' D. Monsecour & K. Monsecour, 2018
* '' Harpa major'' Röding, 1798
* '' Harpa queenslandica'' Berschauer & Petuch, 2016

;Species brought into synonymy:

* ''Harpa conoidalis'' Lamarck, 1822
* ''Harpa crassa'' Krauss, 1848
* ''Harpa fulvomichaelensis'' Orga, 1999
* ''Harpa kawamurai'' Habe, 1970
* ''Harpa laetifica'' Melvill, 1916
* ''Harpa ligata'' Menke, 1828
* ''Harpa minor'' Lamarck, 1822
* ''Harpa multicostata'' G. B. Sowerby I, 1822
* ''Harpa nablium'' Mörch, 1852
* † '' Harpa neozelanica'' Suter, 1917
* ''Harpa nobilis'' Lamarck, 1822
* † '' Harpa pulligera'' Tate, 1889
* ''Harpa robusta'' Röding, 1798
* ''Harpa rosea'' Lamarck, 1816
* ''Harpa striata]'' Lamarck, 1816
* ''Harpa ventricosa'' Lamarck, 1816
* ''Harpa ventricosa'' Lamarck, 1801

;Nomen dubium:
* ''Harpa urniformis'' Perry, 1811

Image:Harpa amouretta 01.JPG, '' Harpa amouretta''
Image:Harpa articularis 01.JPG, '' Harpa articularis''
Image:Hventricosawithshell.png, '' Harpa ventricosa''
Image:Harpa davidis 01.JPG, '' Harpa davidis''
Image:Harpa harpa 01.jpg, ''Harpa harpa''
Image:Harpa doris (2763438292).jpg, '' Harpa doris''
Image:Prosobranchia Trio.jpg, '' Harpa major''

References

* Röding, P.F. 1798. ''Museum Boltenianum sive Catalogus cimeliorum e tribus regnis naturae quae olim collegerat Joa''. Hamburg : Trappii 199 pp.
* Wilson, B. 1994. ''Australian Marine Shells. Prosobranch Gastropods.'' Kallaroo, WA : Odyssey Publishing Vol. 2 370 pp.
* Poppe G.T., Brulet T. & Dance S.P. (1999). ''The family Harpidae.'' Conchological Iconography. ConchBooks, Hackenheim. 69pp.

External links

Harpa Photo Essay

{{Taxonbar, from=Q146930

Taxa named by Peter Friedrich Röding
Gastropod genera