Harmoniconus
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''Harmoniconus'' is a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of
sea snail Sea snail is a common name for slow-moving marine gastropod molluscs, usually with visible external shells, such as whelk or abalone. They share the taxonomic class Gastropoda with slugs, which are distinguished from snails primarily by the ...
s,
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
, the
cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines co ...
and their allies. In the new classification of the family Conidae by Puillandre N., Duda T.F., Meyer C., Olivera B.M. & Bouchet P. (2015), ''Harmoniconus'' has become a subgenus of ''Conus'': ''Conus (Harmoniconus)'' da Motta, 1991 (type species: ''Conus musicus'' Hwass in Bruguière, 1792) represented as ''
Conus ''Conus'' is a genus of predatory sea snails, or cone snails, marine gastropod mollusks in the family Conidae.Bouchet, P.; Gofas, S. (2015). Conus Linnaeus, 1758. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at ...
'' Thiele, 1929 Puillandre N., Duda T.F., Meyer C., Olivera B.M. & Bouchet P. (2015). ''One, four or 100 genera? A new classification of the cone snails''. Journal of Molluscan Studies. 81: 1-23
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Distinguishing characteristics

The Tucker & Tenorio 2009 taxonomy distinguishes ''Harmoniconus'' from ''Conus'' in the following ways:Tucker J.K. & Tenorio M.J. (2009), Systematic Classification of Recent and Fossil Conoidean Gastropods, ConchBooks, Hankenheim, Germany, 295 pp. * Genus ''Conus'' ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'' Linnaeus, 1758 :: Shell characters (living and fossil species) :::The basic shell shape is conical to elongated conical, has a deep anal notch on the shoulder, a smooth
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and ...
and a small operculum. The shoulder of the shell is usually nodulose and the
protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...
is usually multispiral. Markings often include the presence of tents except for black or white color variants, with the absence of spiral lines of minute tents and textile bars. ::Radular tooth (not known for fossil species) :::The
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
has an elongated anterior section with serrations and a large exposed terminating cusp, a non-obvious waist, blade is either small or absent and has a short barb, and lacks a basal spur. ::Geographical distribution :::These species are found in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region. ::Feeding habits :::These species eat other
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s including
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines conn ...
. * Subgenus ''Harmoniconus'' da Motta, 1991 ::Shell characters (living and fossil species) :::The shell is turgid in shape with a low domed spire. The
protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...
is multispiral. The anal notch is shallow. The shell does not have an anterior notch. The shell is either smooth or ornamented with rows of pustules. The
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and ...
is smooth, and the operculum is moderate to large. ::Radular tooth (not known for fossil species) :::The anterior section of the radular tooth is shorter than the length of the posterior section, and the blade is long and covers the length of the anterior section. A basal spur is present, the barb is short, and the posterior fold and/or terminating cusp are partially external. ::Geographical distribution :::The species in this genus occur in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region, and one species (''H. nux'') occurs in the Eastern Pacific region. ::Feeding habits :::These cone snails are vermivorous, meaning that the cones prey on
polychaete Polychaeta () is a paraphyletic class (biology), class of generally marine invertebrate, marine annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that ...
,
Eunicidae Eunicidae is a family of marine polychaetes (bristle worms). The family comprises marine annelids distributed in diverse benthic habitats across Oceania, Europe, South America, North America, Asia and Africa. The Eunicid anatomy typically co ...
and
Nereididae Nereididae (formerly spelled Nereidae) are a family of polychaete worms. It contains about 500 – mostly marine – species grouped into 42 genera. They may be commonly called ragworms or clam worms. Characteristics The prostomium of ...
worms.


Species list

This list of species is based on the information in the
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
(
WoRMS Worms may refer to: *Worm, an invertebrate animal with a tube-like body and no limbs Places *Worms, Germany, a city **Worms (electoral district) *Worms, Nebraska, U.S. *Worms im Veltlintal, the German name for Bormio, Italy Arts and entertainme ...
) list. Species within the genus ''Harmoniconus'' include: The following species names are recognized as "alternate representations" (see full explanation below) in contrast to the traditional system, which uses the genus ''
Conus ''Conus'' is a genus of predatory sea snails, or cone snails, marine gastropod mollusks in the family Conidae.Bouchet, P.; Gofas, S. (2015). Conus Linnaeus, 1758. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at ...
'' for all species in the family: * ''Harmoniconus mcbridei'' (Lorenz, 2005): synonym of '' Conus mcbridei'' Lorenz, 2005 * ''Harmoniconus musicus'' (Hwass in Bruguière, 1792): synonym of '' Conus musicus'' Hwass in Bruguière, 1792 * ''Harmoniconus nanus'' (G.B. Sowerby I, 1833): synonym of ''
Conus nanus ''Conus nanus'' is a species of sea snail, a marine gastropod mollusk in the family (biology), family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conus nanus G. B. Sowerby I, 1833. In: MolluscaBase (2015). Accessed through: Wor ...
'' G.B. Sowerby I, 1833 * ''Harmoniconus nux'' (Broderip, 1833): synonym of '' Conus nux'' Broderip, 1833 * ''Harmoniconus parvatus'' (Walls, 1979): synonym of '' Conus parvatus'' Walls, 1979 * ''Harmoniconus paukstisi'' Tucker, Tenorio & Chaney, 2011:Tucker J.K., Tenorio M.J. & Chaney H.W. (2011) A revision of the status of several conoid taxa from the Hawaiian Islands: Description of Darioconus levieni n. sp., Pionoconus striatus oahuensis n. ssp. and Harmoniconus paukstisi n. sp. (Gastropoda, Conidae). In: Severns M., Shells of the Hawaiian Islands - The Sea Shells: 501-514. Conchbooks, Hackenheim. synonym of ''Conus (Harmoniconus) paukstisi'' (Tucker, Tenorio & Chaney, 2011) represented as '' Conus paukstisi'' (Tucker, Tenorio & Chaney, 2011) * ''Harmoniconus sponsalis'' (Hwass in Bruguière, 1792): synonym of '' Conus sponsalis'' Hwass in Bruguière, 1792


References


Further reading

* Kohn A. A. (1992). "Chronological Taxonomy of ''Conus'', 1758-1840". Smithsonian Institution Press, Washington and London. * Monteiro A. (ed.) (2007)
The Cone Collector 1
1-28. * Berschauer D. (2010). ''Technology and the Fall of the Mono-Generic Family'
The Cone Collector 15
pp. 51–54 * Puillandre N., Meyer C.P., Bouchet P., and Olivera B.M. (2011), ''Genetic divergence and geographical variation in the deep-water Conus orbignyi complex (Mollusca: Conoidea)'', Zoologica Scripta 40(4) 350–363.


External links


Gastropods.com: ''Conidae'' setting forth the genera recognized therein.
{{Taxonbar, from=Q5659331 Conidae Gastropod subgenera