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Haplogroup I-Z63, also known as I1a3 per the International Society of Genetic Genealogy ('ISOGG), is a
Y chromosome The Y chromosome is one of two sex chromosomes (allosomes) in therian mammals, including humans, and many other animals. The other is the X chromosome. Y is normally the sex-determining chromosome in many species, since it is the presence or abse ...
haplogroup. It is correlated with a DYS456 value inferior to 15, but there are exceptions. I-Z63 is most common in
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
,
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
and
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
. Its progenitor is assumed to have lived in
Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
at around 2500 BCE. Within Fennoscandia, I-Z63 has a particularly strong association with Finland. To date, ancient I-Z63 has been found archeologically in the Jutland area of Schleswig, Germany along with Poland and
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
.


Origins

On the basis of analysing samples of volunteers in YDNA sequencing, the YDNA analysis compan
YFull
estimated that I-Z63 formed 4,600 years ago (
2600 BC The 26th century BC was a century that lasted from the year 2600 BC to 2501 BC. Events Crete * c. 2600–2400 BC: Early Minoan I period in Crete. Egypt * c. 2551–2526 BC: Reign of Khufu, second pharaoh of the Fourth Dynasty. The height of the ...
) (95% CI 5,100 <-> 4,000
ybp Before Present (BP) years, or "years before present", is a time scale used mainly in archaeology, geology and other scientific disciplines to specify when events occurred relative to the origin of practical radiocarbon dating in the 1950s. Becaus ...
) with a
TMRCA In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are Common descent, ...
(Time to Most Recent Common Ancestor) of 4,400 years (95% CI 4,900 <-> 3,900
ybp Before Present (BP) years, or "years before present", is a time scale used mainly in archaeology, geology and other scientific disciplines to specify when events occurred relative to the origin of practical radiocarbon dating in the 1950s. Becaus ...
) before present. Geographically I-Z63 is believed to have arisen in or near what is now
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...
(based in part on the current distribution of this haplogroup). The current distribution of I-Z63 shows that there is a very high concentration of I-Z63 on the British Isles. At the same time, the archeological record presents a strong association of I-Z63 to the Wielbark culture and by extension with the
Goths The Goths ( got, 𐌲𐌿𐍄𐌸𐌹𐌿𐌳𐌰, translit=''Gutþiuda''; la, Gothi, grc-gre, Γότθοι, Gótthoi) were a Germanic people who played a major role in the fall of the Western Roman Empire and the emergence of medieval Europe ...
. There is a proposed link between the Goths and British migration, the so-called "Jutish Hypothesis". The "Jutish hypothesis" claims that the Jutes may be synonymous with the
Geats The Geats ( ; ang, gēatas ; non, gautar ; sv, götar ), sometimes called ''Goths'', were a large North Germanic tribe who inhabited ("land of the Geats") in modern southern Sweden from antiquity until the late Middle Ages. They are one of th ...
of southern
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
or their neighbours, the
Gutes The Gutes (old west norse ''Gotar'', old gutnish ''Gutar'') were a North Germanic tribe inhabiting the island of Gotland. The ethnonym is related to that of the ''Goths'' (''Gutans''), and both names were originally Proto-Germanic *''Gutaniz''. ...
. The evidence adduced for this theory includes: * primary sources referring to the Geats (''Geátas'') by alternative names such as ''Iútan'', ''Iótas'' and ''Eotas''; * Asser in his ''Life of Alfred'' (893) identifies the Jutes with the Goths (in a passage claiming that
Alfred the Great Alfred the Great (alt. Ælfred 848/849 – 26 October 899) was King of the West Saxons from 871 to 886, and King of the Anglo-Saxons from 886 until his death in 899. He was the youngest son of King Æthelwulf and his first wife Osburh, who bot ...
was descended, through his mother,
Osburga Osburh or Osburga (also Osburga Oslacsdotter) was the first wife of King Æthelwulf of Wessex and mother of King Alfred the Great. Alfred's biographer, Asser, described her as "a most religious woman, noble in character and noble by birth". ...
, from the ruling dynasty of the Jutish kingdom of
Wihtwara The Wihtwara were one of the tribes of Anglo-Saxon Britain. Their territory was a tribal kingdom located on the Isle of Wight before it was conquered by the Kingdom of Wessex in the late seventh century. The tribe's name is preserved in the na ...
, on the Isle of Wight), and; * the ''
Gutasaga Gutasaga (''Gutasagan'') is a saga regarding the history of Gotland before its Christianization. It was recorded in the 13th century and survives in only a single manuscript, the Codex Holm. B 64, dating to , kept at the National Library of Sweden ...
'' (13th Century) states that some inhabitants of
Gotland Gotland (, ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a province, county, municipality, and diocese. The province includes the islands of Fårö and Gotska Sandön to the ...
left for
mainland Europe Continental Europe or mainland Europe is the contiguous continent of Europe, excluding its surrounding islands. It can also be referred to ambiguously as the European continent, – which can conversely mean the whole of Europe – and, by ...
; large burial sites attributable to either Goths or
Gepids The Gepids, ( la, Gepidae, Gipedae, grc, Γήπαιδες) were an East Germanic tribe who lived in the area of modern Romania, Hungary and Serbia, roughly between the Tisza, Sava and Carpathian Mountains. They were said to share the religion a ...
were found in the 19th century near Willenberg, Prussia (after 1945 Wielbark in Poland). The Jutes invaded and settled in southern Britain in the late
4th century The 4th century (per the Julian calendar and Anno Domini/Common era) was the time period which lasted from 301 (Roman numerals, CCCI) through 400 (Roman numerals, CD). In the West, the early part of the century was shaped by Constantine the Grea ...
during the
Age of Migrations The Migration Period was a period in European history marked by large-scale migrations that saw the fall of the Western Roman Empire and subsequent settlement of its former territories by various tribes, and the establishment of the post-Roman ...
, as part of a larger wave of Germanic settlement in the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
. The Jutish migration to Britain may explain the high concentration of I-Z63 found in modern Britain. However, I-Z63 is notably sparse among modern volunteer testers from Denmark. This is surprising because, in a geographical sense, Denmark encompasses the ancient homeland of the Jutes. Foreign invaders displacing the Jutes from their ancient homeland may explain the relative lack of I-Z63 in Denmark. Even in the year
945 Year 945 ( CMXLV) was a common year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * January 27 – The co-emperors Stephen and Constantine are overthrown barely ...
, the peoples of Jutland were threatened by foreign invaders (yet ironically were posing a threat to other groups elsewhere, such as in England). In 945 King Hacon of Norway arrived in Jutland and slew many of the people there, sending the survivors “far up into the land”. The current distribution of I-Z63 clearly shows that while there is a near absence of I-Z63 in modern Denmark, sizable numbers of I-Z63 men live today in Finland, Norway, Sweden and Aland. Based on the combined evidence, the preferred current working hypothesis puts the progenitor of I-Z63 in ancient
Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
around the year 2000 BCE.


Archeological Record

An individual with the subclade of I-Y21381 was found just south of Denmark in the Schleswig Rathausmarkt excavation in Schleswig, Schleswig-Holstein, Germany. This city on the Jutland peninsula is the successor to the important Viking settlement of
Hedeby Hedeby (, Old Norse ''Heiðabýr'', German language, German ''Haithabu'') was an important Danes, Danish Viking Age (8th to the 11th centuries) trading settlement near the southern end of the Jutland Peninsula, now in the Schleswig-Flensburg dist ...
, which was destroyed in 1066. Sample SWG007 was obtained from grave 179 and dated to 1070-1140 CE. The I-S2077 subclade of haplogroup I-Z63 was sampled on an elite warrior buried in
Bodzia Bodzia is a village in the administrative district of Gmina Lubanie, within Włocławek County, Kuyavian-Pomeranian Voivodeship, in north-central Poland. It lies approximately north-west of Włocławek and south-east of Toruń. Bodzia is also ...
(
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
) in a rich burial from ca. 1010-1020 AD. All artefacts there indicate a strong relation to the Kievian Rus ruling elite, so this man who probably succumbed to combat wounds, was in a close relationship with the Kievian Prince, Sviatopolk the Accursed (son of Vladimir the Great), married to a daughter of the Polish king Boleslaw the Great (Burisleifr from the Scandinavian sagas). The cemetery in Bodzia is exceptional in terms of Scandinavian and Kievian Rus links. The Bodzia man (sample VK157, or burial E864/I) was not a simple warrior from the princely retinue, but he belonged to the princely family himself. His burial is the richest one in the whole cemetery, moreover, Strontium analysis of his teeth enamel shows he was not local. It is assumed that he came to Poland with the Prince of Kiev, Sviatopolk the Accursed, and met a violent death in combat. This corresponds to the events of 1018 AD when Sviatopolk himself disappeared after having retreated from Kiev to Poland. It cannot be excluded that the Bodzia man was Sviatopolk himself, as the genealogy of the
Rurikids The Rurik dynasty ( be, Ру́рыкавічы, Rúrykavichy; russian: Рю́риковичи, Ryúrikovichi, ; uk, Рю́риковичі, Riúrykovychi, ; literally "sons/scions of Rurik"), also known as the Rurikid dynasty or Rurikids, was ...
at this period is extremely sketchy and the dates of birth of many princes of this dynasty may be quite approximative. An autosomal PCA plot of sample VK157 against modern populations has revealed that VK157 matches most closely to the present population of Finland. Taking into account that subclade I-BY316/I-Y7626 bears a very strong association with Finland it may be postulated that I-BY316/I-Y7626 may have been at least partially spread over continental Europe by vikings attached to the Kievan Rus who were originally from Finland. Another archeological sample possibly tied to the Kievan Rus is sample Zeytinliada 14832, which belongs to an 25 to 35 year old adult who lived between 600 and 1000 CE during the Medieval Age and was found in the region now known as Zeytinliada, Erdek, Turkey. This sample has been tied to subclade I-BY316. The genetic profile of this individual does not resemble northern Europeans, so he might be the descendant of a northern European (e.g. part of the Varangian guard of Byzantium) who intermarried with the local population, although a more proximal origin is also possible, as this lineage was found also in Langobards from Hungary. The Rus' provided the earliest members of the Varangian Guard. They were in Byzantine service from as early as 874. Hence it appears plausible that sample Bodzia 157 and Zeytinliada 148 are both connected to the Kievan Rus. I-Z63 has been traced to the Kowalewko burial site in Poland which dates to the Roman Iron Age. In 2017 Polish researchers could successfully assign YDNA haplogroups to 16 individuals who were buried at the site. Out of these 16 individuals three belonged to haplogroup I-Z63, and in particular subclade I-L1237. The Kowalewko archeological site has been associated with the Wielbark culture. The Wielbark culture in turn has been associated with the
Goths The Goths ( got, 𐌲𐌿𐍄𐌸𐌹𐌿𐌳𐌰, translit=''Gutþiuda''; la, Gothi, grc-gre, Γότθοι, Gótthoi) were a Germanic people who played a major role in the fall of the Western Roman Empire and the emergence of medieval Europe ...
. Therefore the subclade I-L1237 of I-Z63 may be seen somewhat as a genetic indicator of the
Gothic Gothic or Gothics may refer to: People and languages *Goths or Gothic people, the ethnonym of a group of East Germanic tribes **Gothic language, an extinct East Germanic language spoken by the Goths **Crimean Gothic, the Gothic language spoken b ...
tribe of
late antiquity Late antiquity is the time of transition from classical antiquity to the Middle Ages, generally spanning the 3rd–7th century in Europe and adjacent areas bordering the Mediterranean Basin. The popularization of this periodization in English ha ...
. There is an academic theory that the Gothic tribe is connected to British migration through the so-called "Jutish Hypothesis", which would explain why I-L1237 is so strongly associated both with British migration and with Gothic migration patterns. I-Z63 was found in a late 6th Century cemetery in Collegno, Italy, near the city of Torino. The Collegno burial site is associated with Gothic and Lombard remains and dated to the late 6th Century. The remains belong to I-FT104588, which is a subclade of I-BY316/I-Y7626. This discovery adds another historical connection of I-Z63 to the Gothic migrations of the early Medieval Period. Another I-Z63 archeological find at the Crypta Balbi site in Rome was dated to Late Antiquity (400-600 CE) and reported to be Y-DNA haplogroup I-Y7234. Interestingly, Lombard-associated ornaments have been excavated at this site, pointing to connections with central Europe. Additionally, five of the seven individuals from this site, including the I-Z63 individual, were classified by ChromoPainter into a cluster with more haplotype sharing with central/northern Europeans. Excavated in Hungary, Tiszafüred 798 was a man, associated with the Late Avar cultural group, who lived between 700 and 800 CE during the Medieval Age and was found in the region now known as Majoros-halom, Tiszafüred. Likewise Tiszafüred 509 was a man who lived between 800 and 830 CE. Both were matched to haplogroup I-S10360. Avars, under the rule of a khagan called Bayan, 565 AD, allied with the Longobards and defeated the Germanic Gepides in the Danube Plains. In 7th c. AD they continued raiding Western Europe. They helped the lombard king Grimoald to vanquish a rebellion in Frioul, 663 AD. Hence a plausible explanation exists how a haplogroup which is strongly associated with the Gothic migrations is also attached to the Avar cultural group. Also found in Hungary was Magyarhomorog 88 who belongs to subclade I-PR683, he was a man who lived between 1000 and 1100 CE during the Medieval Age and was found in the region now known as Kónyadomb, Magyarhomorog, Hungary.


Significant Subclades

Even though I-Z63 itself is not considered a Nordic haplogroup, it does with I-BY316/I-Y7626 contain a significant Nordic subclade, whereby I-BY316 is heavily weighted towards Finland, Sweden, Norway, Iceland and Aland.


Prominent members of I-Z63

Sviatopolk the Accursed (son of Vladimir the Great) is believed to have carried the I-S2077 subclade of Z63. By patrilineal extension this may also make
Rurik Rurik (also Ryurik; orv, Рюрикъ, Rjurikŭ, from Old Norse '' Hrøríkʀ''; russian: Рюрик; died 879); be, Рурык, Ruryk was a semi-legendary Varangian chieftain of the Rus' who in the year 862 was invited to reign in Novgoro ...
an I-Z63.
Governor William Bradford William Bradford ( 19 March 15909 May 1657) was an English Puritan separatist originally from the West Riding of Yorkshire in Northern England. He moved to Leiden in Holland in order to escape persecution from King James I of England, and then ...
,
Mayflower ''Mayflower'' was an English ship that transported a group of English families, known today as the Pilgrims, from England to the New World in 1620. After a grueling 10 weeks at sea, ''Mayflower'', with 102 passengers and a crew of about 30, r ...
passenger and leader of the Plymouth Colony. At least two reported descendants have tested under I-Z63 to I-Y21381 > Y21371 > Y21370 > Y21372 > FGC72882.


References

{{reflist Human Y-DNA haplogroups Human evolution Population genetics Classical genetics