''Halichoeres richmondi'', commonly called the Richmond's wrasse or chain-lined wrasse, is a fish species in the

wrasse The wrasses are a family, Labridae, of marine fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into 9 subgroups or tribes. They are typically small, most of them le ...

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

native from the central

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

Description

The Richmond's wrasse is a small fish that can reach a maximum length of 19 cm. It has a thin, elongate body with a terminal mouth and a more pointed snout than other wrasse belonging to genus Halichoeres. Body coloration has few variations according to age. The juvenile has a light blue-green background color with many orange stripes. It has also two black

ocellus A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l ...

: the first one on top part of its

caudal peduncle Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

and the second one in the middle of its

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

. A black spot occurs on the first rays of the

. Juveniles and females have in common an orange

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

. Mature male are quite different. Their body is greenish with many fin blue lines that are chain-like. A concave head profile with a green-brown color. The external border of their dorsal, caudal and anal fins are blue.

Distribution & habitat

The chain-lined wrasse is widespread throughout the tropical and subtropical waters of the central

, from

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

and from

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...

to south

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

. This wrasse occurs in shallow waters from lagoons and channels rich in soft coral down to a depth of 12 meters.Lieske, E. and R. Myers, 1994. Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Harper Collins Publishers, 400 p.Kuiter, R.H. and T. Tonozuka, 2001. Pictorial guide to Indonesian reef fishes. Part 2. Fusiliers - Dragonets, Caesionidae - Callionymidae. Zoonetics, Australia. 304-622 p.Rudie Kuiter, "Labridae fishes: wrasses", Aquatic Photographics, 2010,

Biology

The chain-lined wrasse lives and feeds in small loose groups. It is a

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

predator that feeds mainly on small marine

invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

such as

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil sp ...

, worms and

echinoderms An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea li ...

captured on or in the substrate. Like most wrasse, the chain-lined wrasse is a

protogynous hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...

, i.e. individuals start life as females with the capability of turning male later on.

Conservation status

The species is targeted but not thought to be threatened by the aquarium trade. It is listed as Least Concern (LC) on the IUCN.

Species description and etymology

''Halichoeres richmondi'' was described in 1928 by the

American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...

Henry Weed Fowler Henry Weed Fowler (March 23, 1878 – June 21, 1965) was an American zoologist born in Holmesburg, Pennsylvania. He studied at Stanford University under David Starr Jordan. He joined the Academy of Natural Sciences in Philadelphia and worked as ...

and

Barton Appler Bean Barton Appler Bean was an American ichthyologist, born May 21, 1860 in Lancaster County, Pennsylvania and died June 16, 1947 in Chemung, New York, after falling from a bridge. He was the brother of the ichthyologist Tarleton Hoffman Bean (1846-1 ...

with the type locality given as Inamucan Bay on

Mindanao Mindanao ( ) ( Jawi: مينداناو) is the second-largest island in the Philippines, after Luzon, and seventh-most populous island in the world. Located in the southern region of the archipelago, the island is part of an island group of ...

in the Philippines. The specific name honours the

ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...

Charles Wallace Richmond Charles Wallace Richmond (December 31, 1868 – May 19, 1932) was an American ornithologist. He is best remembered for a compilation of the Latin names of birds that is called the Richmond Index. Life and work He was born in Kenosha, Wisconsin an ...

(1868-1932) of the

United States National Museum The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums and education and research centers, the largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge". Founded ...

References

External links

* {{Taxonbar, from=Q2606636 richmondi Taxa named by Henry Weed Fowler Taxa named by Barton Appler Bean Fish described in 1928