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''Hagenia'' is a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
with the sole species ''Hagenia abyssinica'', native to the high-elevation
Afromontane The Afromontane regions are subregions of the Afrotropical realm, one of the Earth's eight biogeographic realms, covering the plant and animal species found in the mountains of Africa and the southern Arabian Peninsula. The Afromontane regions ...
regions of central and eastern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. It also has a disjunct distribution in the high mountains of East Africa from
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
and
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
in the north, through
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
,
Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...
,
Burundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili language, Swahili: ''Jamuhuri ya Burundi''; French language, French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the ...
,
Democratic Republic of Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
, and
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
, to
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeast ...
and
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...
in the south. A member of the
rose family Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are '' Alchemilla'' (270), ''Sorbus ...
, its closest relative is the Afromontane genus '' Leucosidea''.


Nomenclature

It is known in
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...
as African redwood, East African rosewood, brayera, cusso, hagenia, or kousso, in
Amharic Amharic ( or ; (Amharic: ), ', ) is an Ethiopian Semitic language, which is a subgrouping within the Semitic branch of the Afroasiatic languages. It is spoken as a first language by the Amharas, and also serves as a lingua franca for all oth ...
as ''kosso'', and in Swahili as ''mdobore'' or ''mlozilozi''.
Synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of the species include ''Banksia abyssinica'', ''Brayera anthelmintica'', ''Hagenia abyssinica'' var. ''viridifolia'' and ''Hagenia anthelmintica''.


Description

It is a
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
up to 20 m in height, with a short trunk, thick branches, and thick, peeling bark. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are up to 40 cm long, compound with 7-13 leaflets, each leaflet about 10 cm long with a finely serrated margin, green above, silvery-haired below. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are white to orange-buff or pinkish-red, produced in
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s 30–60 cm long.


Distribution

It is generally found from 2000–3000 m elevation, in areas receiving 1000–1500 mm of rainfall annually. It can be found growing in mixed afromontane forest with ''
Podocarpus ''Podocarpus'' () is a genus of conifers, the most numerous and widely distributed of the podocarp family, the Podocarpaceae. The name comes from Greek πούς (poús, “foot”) + καρπός (karpós, “fruit”). ''Podocarpus'' species ...
'', ''
Afrocarpus ''Afrocarpus'' is a genus of conifers of the family Podocarpaceae. Two to six species are recognized. They are evergreen trees native to Africa. ''Afrocarpus'' was designated a genus in 1989, when several species formerly classified in ''Podocarp ...
'', and other trees, and in drier afromontane forests and woodlands where ''Hagenia'' is dominant, or in mixed stands of ''Hagenia'' and ''
Juniperus procera ''Juniperus procera'' (known by the common English names African juniper, African pencil-cedar, East African juniper, East African-cedar, and Kenya-cedar) is a coniferous tree native to mountainous areas in Africa and the Arabian Peninsula. It is ...
''. It is often found near the upper limit of forest growth, giving way to giant heather zones above it.


Ecology

''Hagenia'' is used as a food plant by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including
turnip moth ''Agrotis segetum'', sometimes known as the turnip moth, is a moth of the family Noctuidae. The species was first described by Michael Denis and Ignaz Schiffermüller in 1775. It is a common European species and it is found in Africa and across ...
.


References


External links


''Hagenia abyssinica'' at the AgroForestry Tree Database (World Agroforestry Centre)"The Kosso Tree"
is a watercolor by William Simpson from 1868 * {{Taxonbar, from1=Q13108105, from2=Q1785081 Agrimoniinae Afromontane flora Trees of Africa Trees of the Democratic Republic of the Congo Trees of Ethiopia Medicinal plants of Africa Monotypic Rosaceae genera Plant dyes Taxa named by Johann Friedrich Gmelin